I have been assured by several people, that there are no coral-reefs on the west coast of Africa,[4] or round the islands in the Gulf of Guinea. This perhaps may be attributed, in part, to the sediment brought down by the many rivers debouching on that coast, and to the extensive mud-banks, which line great part of it. But the islands of St. Helena, Ascension, the Cape Verdes, St. Paul’s, and Fernando Noronha, are, also, entirely without reefs, although they lie far out at sea, are composed of the same ancient volcanic rocks, and have the same general form, with those islands in the Pacific, the shores of which are surrounded by gigantic walls of coral-rock. With the exception of Bermuda, there is not a single coral-reef in the central expanse of the Atlantic Ocean. It will, perhaps, be suggested that the quantity of carbonate of lime in different parts of the sea, may regulate the presence of reefs. But this cannot be the case, for at Ascension, the waves charged to excess precipitate a thick layer of calcareous matter on the tidal rocks; and at St. Jago, in the Cape Verdes, carbonate of lime not only is abundant on the shores, but it forms the chief part of some upraised post-tertiary strata. The apparently capricious distribution, therefore, of coral-reefs, cannot be explained by any of these obvious causes; but as the study of the terrestrial and better known half of the world must convince every one that no station capable of supporting life is lost,—nay more, that there is a struggle for each station, between the different orders of nature,—we may conclude that in those parts of the intertropical sea, in which there are no coral-reefs, there are other organic bodies supplying the place of the reef-building polypifers. It has been shown in the chapter on Keeling atoll that there are some species of large fish, and the whole tribe of Holothuriæ which prey on the tenderer parts of the corals. On the other hand, the polypifers in their turn must prey on some other organic beings; the decrease of which from any cause would cause a proportionate destruction of the living coral. The relations, therefore, which determine the formation of reefs on any shore, by the vigorous growth of the efficient kinds of coral, must be very complex, and with our imperfect knowledge quite inexplicable. From these considerations, we may infer that changes in the condition of the sea, not obvious to our senses, might destroy all the coral-reefs in one area, and cause them to appear in another: thus, the Pacific or Indian Ocean might become as barren of coral-reefs as the Atlantic now is, without our being able to assign any adequate cause for such a change.