_Section III_—ON THE DEPTHS AT WHICH REEF-BUILDING POLYPIFERS CAN LIVE I have already described in detail, which might have appeared trivial, the nature of the bottom of the sea immediately surrounding Keeling atoll; and I will now describe with almost equal care the soundings off the fringing-reefs of Mauritius. I have preferred this arrangement, for the sake of grouping together facts of a similar nature. I sounded with the wide bell-shaped lead which Captain Fitzroy used at Keeling Island, but my examination of the bottom was confined to a few miles of coast (between Port Louis and Tomb Bay) on the leeward side of the island. The edge of the reef is formed of great shapeless masses of branching Madrepores, which chiefly consist of two species,—apparently _M. corymbosa_ and _ pocillifera_,—mingled with a few other kinds of coral. These masses are separated from each other by the most irregular gullies and cavities, into which the lead sinks many feet. Outside this irregular border of Madrepores, the water deepens gradually to twenty fathoms, which depth generally is found at the distance of from half to three-quarters of a mile from the reef. A little further out the depth is thirty fathoms, and thence the bank slopes rapidly into the depths of the ocean. This inclination is very gentle compared with that outside Keeling and other atolls, but compared with most coasts it is steep. The water was so clear outside the reef, that I could distinguish every object forming the rugged bottom. In this part, and to a depth of eight fathoms, I sounded repeatedly, and at each cast pounded the bottom with the broad lead, nevertheless the arming invariably came up perfectly clean, but deeply indented. From eight to fifteen fathoms a little calcareous sand was occasionally brought up, but more frequently the arming was simply indented. In all this space the two Madrepores above mentioned, and two species of Astræa, with rather large[30] stars, seemed the commonest kinds; and it must be noticed that twice at the depth of fifteen fathoms, the arming was marked with a clean impression of an Astræa. Besides these lithophytes, some fragments of the _Millepora alcicornis,_ which occurs in the same relative position at Keeling Island, were brought up; and in the deeper parts there were large beds of a Seriatopora, different from _S. subulata_, but closely allied to it. On the beach within the reef, the rolled fragments consisted chiefly of the corals just mentioned, and of a massive Porites, like that at Keeling atoll, of a Meandrina, _ Pocillopora verrucosa_, and of numerous fragments of Nullipora. From fifteen to twenty fathoms the bottom was, with few exceptions, either formed of sand, or thickly covered with Seriatopora: this delicate coral seems to form at these depths extensive beds unmingled with any other kind. At twenty fathoms, one sounding brought up a fragment of Madrepora apparently _M.