energy borrowed from the sun, and on the other hand to expend it, in a
discontinuous and explosive way, in movements of locomotion. Even
to-day, perhaps, a chlorophyl-bearing Infusorian such as the Euglena may
symbolize this primordial tendency of life, though in a mean form,
incapable of evolving. Is the divergent development of the two kingdoms
related to what one may call the oblivion of each kingdom as regards one
of the two halves of the programme? Or rather, which is more likely, was
the very nature of the matter, that life found confronting it on our
planet, opposed to the possibility of the two tendencies evolving very
far together in the same organism? What is certain is that the vegetable
has trended principally in the first direction and the animal in the
second. But if, from the very first, in making the explosive, nature had
for object the explosion, then it is the evolution of the animal, rather
than that of the vegetable, that indicates, on the whole, the
fundamental direction of life.
The "harmony" of the two kingdoms, the complementary characters they
display, might then be due to the fact that they develop two tendencies
which at first were fused in one. The more the single original tendency
grows, the harder it finds it to keep united in the same living being
those two elements which in the rudimentary state implied each other.
Hence a parting in two, hence two divergent evolutions; hence also two
series of characters opposed in certain points, complementary in others,
but, whether opposed or complementary, always preserving an appearance
of kinship. While the animal evolved, not without accidents along the
way, toward a freer and freer expenditure of discontinuous energy, the
plant perfected rather its system of accumulation without moving. We
shall not dwell on this second point. Suffice it to say that the plant
must have been greatly benefited, in its turn, by a new division,
analogous to that between plants and animals. While the primitive
vegetable cell had to fix by itself both its carbon and its nitrogen, it
became able almost to give up the second of these two functions as soon
as microscopic vegetables came forward which leaned in this direction
exclusively, and even specialized diversely in this still complicated
business. The microbes that fix the nitrogen of the air and those which
convert the ammoniacal compounds into nitrous ones, and these again into
nitrates, have, by the same splitting up of a tendency primitively one,
rendered to the whole vegetable world the same kind of service as the
vegetables in general have rendered to animals. If a special kingdom
were to be made for these microscopic vegetables, it might be said that
in the microbes of the soil, the vegetables and the animals, we have
before us the _analysis_, carried out by the matter that life found at
its disposal on our planet, of all that life contained, at the outset,
in a state of reciprocal implication. Is this, properly speaking, a
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