But when we turn to the higher _Vertebrata_, the results of recent
investigations, however we may sift and criticise them, seem to me to
leave a clear balance in favour of the doctrine of the evolution
of living forms one from another. Nevertheless, in discussing this
question, it is very necessary to discriminate carefully between the
different kinds of evidence from fossil remains which are brought
forward in favour of evolution.
Every fossil which takes an intermediate place between forms of
life already known, may be said, so far as it is intermediate, to be
evidence in favour of evolution, inasmuch as it shows a possible road
by which evolution may have taken place. But the mere discovery of
such a form does not, in itself, prove that evolution took place by
and through it, nor does it constitute more than presumptive evidence
in favour of evolution in general. Suppose A, B, C to be three
forms, while B is intermediate in structure between A and C. Then the
doctrine of evolution offers four possible alternatives. A may have
become C by way of B; or C may have become A by way of B; or A and
C may be independent modifications of B; or A, B, and C may be
independent modifications of some unknown D. Take the case of the
Pigs, the _Anoplotheridae_, and the Ruminants. The _Anoplotheridae_
are intermediate between the first and the last; but this does not
tell us whether the Ruminants have come from the Pigs, or the Pigs
from Ruminants, or both from _Anoplotheridae_, or whether Pigs,
Ruminants, and _Anoplotheridae_ alike may not have diverged from some
common stock.
But if it can be shown that A, B, and C exhibit successive stages in
the degree of modification, or specialization, of the same type; and
if, further, it can be proved that they occur in successively
newer deposits. A being in the oldest and C in the newest, then the
intermediate character of B has quite another importance, and I should
accept it, without hesitation, as a link in the genealogy of C. I
should consider the burden of proof to be thrown upon anyone who
denied C to have been derived from A by way of B, or in some closely
analogous fashion; for it is always probable that one may not hit upon
the exact line of filiation, and, in dealing with fossils, may mistake
uncles and nephews for fathers and sons.
I think it necessary to distinguish between the former and the latter
classes of intermediate forms, as _intercalary types_ and _linear
types_. When I apply the former term, I merely mean to say that as
a matter of fact, the form B, so named, is intermediate between the
others, in the sense in which the _Anoplotherium_ is intermediate
between the Pigs and the Ruminants--without either affirming, or
denying, any direct genetic relation between the three forms involved.
When I apply the latter term, on the other hand, I mean to express the
opinion that the forms A, B, and C constitute a line of descent, and
that B is thus part of the lineage of C.
Public-domain text, read in full here on John Shaqi.
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