Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
Comparatively few, then, though the facts may be of which I have given
some examples, in my opinion they are amply sufficient for the purpose
in hand. This purpose is to show that the question which we are now
considering is very far from being a closed question; and, therefore,
that the school of Weismann is much too precipitate in alleging that
there is neither any necessity for, nor evidence of, the so-called
Lamarckian factors[85]. And this opinion, whatever it may be worth, is
at all events both deliberate and impartial. As one of the first to
doubt the transmission of acquired characters, and as one who has spent
many years in experimental inquiries upon the subject, any bias that I
may have is assuredly against the Lamarckian principles--seeing that
nearly all my experiments have yielded negative results. It was Darwin
himself who checked this bias. But if the ultra-Darwinians of the last
ten years had succeeded in showing that Darwin was mistaken, I should be
extremely glad to fall into line with them. As already shown, however,
they have in no way affected this question as it was left by Galton in
1875. And if it be supposed a matter of but little importance whether we
agree with Galton in largely diminishing the comparative potency of the
Lamarckian principles, or whether we agree with Weismann in abolishing
them together, it cannot be too often repeated that such is an entirely
erroneous view. No matter how faintly or how fitfully acquired
characters may be transmitted, in so far as they are likewise adaptive
characters, their transmission (and therefore their development) must be
cumulative. Hence, the only effect of attenuating our estimate of their
_intensity_, is that of increasing our estimate of their
_duration_--i.e. of the time over which they have to operate in order to
produce important results. And, even so, it is to be remembered that
the importance of such results is not to be estimated by the magnitude
of modification. Far more is it to be estimated by the character of
modification as adaptive. For if functionally produced changes, and
changes produced in adaptive response to the environment, are ever
transmitted in a cumulative manner, a time must sooner or later arrive
when they will reach a selective value in the struggle for
existence--when, of course, they will be rapidly augmented by natural
selection. Thus, if in any degree operative at all, the great function
of these principles must be that of supplying to natural selection those
incipient stages of adaptive modifications in all cases where, but for
their agency, there would have been nothing of the kind to select.
Themselves in no way dependent on adaptive modifications having already
attained a selective value, these Lamarckian principles are (under the
Darwinian theory) direct causes of determinate variation in adaptive
lines; and variation in those lines being cumulative, the result is that
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account