Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
* * * * *
I have dwelt thus at some length upon a mere matter of definition
because, as we shall now find, although it is but a matter of
definition, it is fraught with consequences of no small importance to
the general theory of descent. Starting from an erroneous definition of
the theory of natural selection as primarily a theory of the origin of
species, both friends and foes of the theory have concluded that the
principle of utility must by hypothesis be of universal occurrence so
far as species are concerned; whereas, if once these naturalists were to
perceive that their definition of the theory is erroneous, they would
likewise perceive that their conclusion cannot follow deductively from
the theory itself. If such a conclusion is to be established at all, it
can only be by other and independent evidence of the inductive kind--to
wit, by actual observation.
Hence we see the importance of starting with an accurate definition of
the theory before proceeding to examine the doctrine of utility as of
universal application to species--a doctrine which, as just stated, has
been habitually and expressly deduced from the theory. This doctrine
occurs in two forms; or, more correctly, there are with reference to
this subject two distinct doctrines, which partly coincide and partly
exclude one another. First, it is held by some naturalists that all
species must necessarily owe their origin to natural selection. And
secondly, it is held by other naturalists, that not only all species,
but likewise all specific characters must necessarily do the same. Let
us consider these two doctrines separately.
The first, and less extensive doctrine, rests on the deduction that
every species must owe its differentiation as a species to the evolution
of at least one adaptive character, which is peculiar to that species.
Although, when thus originated, a species may come to present any
number of other peculiar characters of a non-adaptive kind, these merely
indifferent peculiarities are supposed to hang, as it were, on the peg
supplied by the one adaptive peculiarity; it is the latter which
conditions the species, and so furnishes an opportunity for any number
of the former to supervene. But without the evolution of at least one
adaptive character there could have been no distinct species, and
therefore no merely adventitious characters as belonging to that
species. I will call this the Huxleyan doctrine, because Professor
Huxley is its most express and most authoritative supporter.
Public-domain text, read in full here on John Shaqi.
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