Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
"Elsewhere (_Origin of Species_, p. 158) Mr. Darwin points out that
modifications which appear to present obvious utility are often
found on further examination to be really useless. This latter
consideration, therefore, may be said to act as a foil to the one
against which I am arguing, namely, that modifications which appear
to be useless may nevertheless be useful. But here is a still more
suggestive consideration, also derived from Mr. Darwin's writings.
Among our domesticated productions changes of structure--or even
structures wholly new--not unfrequently arise, which are in every
way analogous to the apparently useless distinctions between wild
species. Take, for example, the following most instructive case:--
[Illustration: Fig. 2.--Old Irish Pig, showing jaw-appendages
(after Richardson).]
"'Another curious anomaly is offered by the appendages described by
M. Eudes-Deslongchamps as often characterizing the Normandy pigs.
These appendages are always attached to the same spot, to the
corners of the jaws; they are cylindrical, about three inches in
length, covered with bristles, and with a pencil of bristles rising
out of a sinus on one side; they have a cartilaginous centre with
two small longitudinal muscles; they occur either symmetrically on
both sides of the face, or on one side alone. Richardson figures
them on the gaunt old Irish Greyhound pig; and Nathusius states
that they occasionally appear in all the long-eared races, but are
not strictly inherited, for they occur or fail in the animals of
the same litter. As no wild pigs are known to have analogous
appendages, we have at present no reason to suppose that their
appearance is due to reversion; and if this be so, we are forced to
admit that a somewhat complex, though apparently useless, structure
may be suddenly developed without the aid of selection[97].'"
[97] _Variation_, &c. vol. i. pp. 78-79.
To this case Mr. Wallace objects:--
"But it is expressly stated that they are not constant; they appear
'frequently' or 'occasionally,' they are 'not strictly inherited,
for they occur or fail in animals of the same litter'; and they are
not always symmetrical, sometimes appearing on one side of the face
alone. Now, whatever may be the cause or explanation of these
anomalous appendages, they cannot be classed with 'specific
characters,' the most essential features of which are, that they
_are_ symmetrical, that they _are_ inherited, and that they _are_
constant[98]."
[98] _Darwinism_, pp. 139-40.
But, to begin with, I have not classed these appendages with "specific
characters," nor maintained that Normandy pigs ought to be regarded as
specifically distinct on account of them. What I said was:--
Public-domain text, read in full here on John Shaqi.
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