Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
Hence, it is plainly _impossible_ to attribute this modification to
natural selection, either as acting directly on the modified parts
themselves, or indirectly through correlation of growth. And as the
modification is of specific magnitude on the one hand, while it presents
all "the most essential features of specific characters" on the other, I
do not see any means whereby Mr. Wallace can meet it on his _a priori_
principles. It would be useless to answer that these characters,
although conforming to all his tests of specific characters, differ in
respect of being deleterious, and would therefore lead to extermination
were the animals in a wholly wild state; because, considered as an
argument, this would involve the assumption that, apart from natural
selection, only deleterious characters can arise under nature--i. e.
that merely "indifferent" characters can never do so, which would be
absurd. Indeed, I have chosen this case of the niata cattle expressly
because their strongly marked peculiarities _are_ deleterious, and
therefore exclude Mr. Wallace's appeal to the argument from ignorance of
a possible utility. But if even these pronounced and deleterious
peculiarities can arise and be perpetuated with such constancy and
fidelity, much more is this likely to be the case with less pronounced
and merely neutral peculiarities.
It may, however, be further objected that these cattle are not
improbably the result of _artificial_ selection. It may be suggested
that the semi-monstrous breed originated in a single congenital
variation, or "sport," which was isolated and multiplied as a curiosity
by the early settlers. But even if such be the explanation of this
particular case, the fact would not weaken our illustration. On the
contrary, it would strengthen our general argument, by showing an
additional means whereby indifferent specific characters can arise and
become fixed in a state of nature. As it seems to me extremely probable
that the niata cattle did originate in a congenital monstrosity, which
was then isolated and multiplied by human agency (as is known to have
been the case with the "ancon sheep"), I will explain why this tends to
strengthen our general argument.
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