Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
[119] _Divergent Evolution through Cumulative Segregation_, Linn.
Journ. Zoology, vol. xx. p. 215.
[120] _Habit and Intelligence_, p. 241.
[121] Allusion may here again be made to the case of the niata
cattle. For here is a case where a very extreme variety is
certainly not unstable, nor produced in varying proportions
from the parent form. Moreover, as we have seen in the
preceding chapter, this almost monstrous variety most probably
originated as an individual sport--being afterwards maintained
and multiplied for a time by artificial selection. Now,
whether or not this was the case, we can very well see that it
may have been. Hence it will serve to illustrate another
possibility touching the origin and maintenance of useless
specific characters. For what is to prevent an individual
congenital variation of any kind (provided it be not harmful)
from perpetuating itself as a "varietal," and eventually,
should offspring become sufficiently numerous, a "specific
character"? There is nothing to prevent this, save panmixia,
or the presence of free intercrossing. But, as we shall see in
the next division of this treatise, there are in nature many
forms of isolation. Hence, as often as a small number of
individuals may have experienced isolation in any of its
forms, opportunity for perpetuation will have been given to
any congenital variations which may happen to arise. Should
any of these be pronounced variations, it would afterwards be
ranked as a specific character. I do not myself think that
this is the way in which indifferent specific characters
_usually_ originate. On the contrary, I believe that their
origin is most frequently due to the influence of isolation on
the average characters of the whole population, as briefly
stated in the text. But here it seems worth while to notice
this possibility of their occasionally arising as merely
individual variations, afterwards perpetuated by any of the
numerous isolating conditions which occur in nature. For, if
this can be the case with a varietal form so extreme as to
border on the monstrous, much more can it be so with such
minute differences as frequently go to constitute specific
distinctions. It is the business of species-makers to search
out such distinctions, no matter how trivial, and to record
them as "specific characters." Consequently, wherever in
nature a congenital variation happens to arise, and to be
perpetuated by the force of heredity alone under any of the
numerous forms of isolation which occur in nature, there will
be a case analogous to that of the niata cattle.
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