Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
First of all, then, what is this distinction? Take, for example, the
case of the Porto Santo rabbits. To almost every naturalist who reads
what has been said touching these animals, it will have appeared that
the connexion in which they are adduced is wholly irrelevant to the
question in debate. For, it will be said that the very fact of the
seemingly specific differentiation of these animals having proved to be
illusory when some of them were restored to their ancestral conditions,
is proof that their peculiar characters are not specific characters; but
only what Mr. Wallace would term "individual characters," or variations
that are not _inherited_. And the same remark applies to all the other
cases which have been adduced to show the generality and extent of
climatic variation, both in other animals and also in plants. Why, then,
it will be asked, commit the absurdity of adducing such cases in the
present discussion? Is it not self-evident that however general, or
however considerable, such merely individual, or non-heritable,
variations may be, they cannot possibly have ever had anything to do
with the origin of _species_? Therefore, is it not simply preposterous
to so much as mention them in relation to the question touching the
utility of specific characters?
Well, whether or not it is absurd and preposterous to consider climatic
variations in connexion with the origin of species, will depend, and
depend exclusively, on what it is that we are to understand by a
species. Hitherto I have assumed, for the sake of argument, that we all
know what is meant by a species. But the time has now come for showing
that such is far from being the case. And as it would be clearly absurd
and preposterous to conclude anything with regard to specific characters
before agreeing upon what we mean by a character as specific, I will
begin by giving all the logically possible definitions of a species.
1. _A group of individuals descended by way of natural generation from
an originally and specially created type._
This definition may be taken as virtually obsolete.
2. _A group of individuals which, while fully fertile_ inter se, _are
sterile with all other individuals--or, at any rate, do not generate
fully fertile hybrids._
This purely physiological definition is not nowadays entertained by any
naturalist. Even though the physiological distinction be allowed to
count for something in otherwise doubtful cases, no systematist would
constitute a species on such grounds alone. Therefore we need not
concern ourselves with this definition, further than to observe that it
is often taken as more or less supplementary to each of the following
definitions.
3. _A group of individuals which, however many characters they share
with other individuals, agree in presenting one or more characters of a
peculiar kind, with some certain degree of distinctness._
Public-domain text, read in full here on John Shaqi.
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