Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
Of course this definition rests upon the dogma of utility as a necessary
attribute of characters _quâ_ specific--i.e. the dogma against which the
whole of the present discussion is directed. Therefore all I need say
with reference to it is, that at any rate it cannot be adduced in any
argument where the validity of its basal dogma is in question. For it
would be a mere begging of this question to argue that every species
must present at least one peculiar and adaptive character, because,
according to definition, unless an organic type does present at least
one such character, it is not a specific type. Moreover, and quite apart
from this, it is to be hoped that naturalists as a body will never
consent to base their diagnostic work on what at best must always be a
highly speculative extension of the Darwinian theory. While, lastly, if
they were to do so with any sort of consistency, the precise adaptation
which each peculiar character subserves, and which because of this
adaptation is constituted a character of specific distinction, would
have to be determined by actual observation. For no criterion of
specific distinction could be more vague and mischievous than this one,
if it were to be applied on grounds of mere inference that such and such
a character, because seemingly constant, must "necessarily" be either
useful, vestigial, or correlated.
Such then, as far as I can see, are all the definitions of a species
that are logically possible[122]. Which of them is chosen by those who
maintain the necessary usefulness of all specific characters? Observe,
it is for those who maintain this doctrine to choose their definition:
it is not for me to do so. My contention is, that the term does not
admit of any definition sufficiently close and constant to serve as a
basis for the doctrine in question--and this for the simple reason that
species-makers have never agreed among themselves upon any criterion of
specific distinction. My opponents, on the other hand, are clearly bound
to take an opposite view, because, unless they suppose that there is
some such definition of a species, they would be self-convicted of the
absurdity of maintaining a great generalization on a confessedly
untenable basis. For example, a few years ago I was allowed to raise a
debate in the Biological Section of the British Association on the
question to which the present chapters are devoted. But the debate ended
as I had anticipated that it must end. No one of the naturalists present
could give even the vaguest definition of what was meant by a
species--or, consequently, of a character as specific. On this account
the debate ended in as complete a destruction as was possible of the
doctrine that all the distinctive characters of every species must
necessarily be useful, vestigial, or correlated. For it became
unquestionable that the same generalization admitted of being made, with
the same degree of effect, touching all the distinctive characters of
every "snark."
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