Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
With regard to experiments, so few have thus far been made upon the
subject, that objections founded on their negative results do not carry
much weight--especially when we remember that these results have not
been uniformly negative, but sometimes positive, as shown in Chapter VI.
With regard to plants and animals in a state of nature, the objection is
wholly futile, for the simple reason that in as many cases as changed
conditions of life may have caused an hereditary change of specific
type, there is now no means of obtaining "evidence" upon the subject.
But we are not on this account entitled to conclude against the
probability of such changes of specific type having been more or less
frequently thus produced. And still less can we be on this account
entitled to conclude against the _possibility_ of such a change having
ever occurred in any single instance. Yet this is what must be concluded
by any one who maintains that the origin of all species--and, _a
fortiori_, of all specific characters--must _necessarily_ have been due
to natural selection.
Now, if all this be admitted--and I do not see how it can be reasonably
questioned--consider how important its bearing becomes on the issue
before us. If germ-plasm (using this term for whatever it is that
constitutes the material basis of heredity) is ever capable of having
its congenital endowments altered by the direct action of external
conditions, the resulting change of hereditary characters, whatever else
it may be, need not be an adaptive change. Indeed, according to
Weismann's theory of germ-plasm, the chances must be infinitely against
the change being an adaptive one. On the theory of pangenesis--that is
to say, on the so-called Lamarckian principles--there would be much more
reason for entertaining the possibly adaptive character of hereditary
change due to the direct action of the environment. Therefore we arrive
at this curious result. The more that we are disposed to accept
Weismann's theory of heredity, and with it the corollary that natural
selection is the sole cause of adaptive modification in species the less
are we entitled to assume that all specific characters must necessarily
be adaptive. Seeing that in nature there are presumably many cases like
those of Hoffmann's plants, Weismann's butterflies, &c., where the
hereditary qualities of germ-plasm have (on his hypothesis) been
modified by changed conditions of life, we are bound to believe that, in
all cases where such changes do not happen to be actively deleterious,
they will persist. And inasmuch as characters which are only of
"specific" value must be the characters most easily--and therefore most
frequently--induced by any slight changes in the constitution of
germ-plasm, while, for the same reason (namely, that of their trivial
nature) they are least likely to prove injurious, it follows that the
less we believe in the functionally-produced adaptations of Lamarck, the
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account