Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
But what, it may well be asked, is the use of thus insisting upon
well-known facts, which nobody will dispute? Well, in the first place,
we have already seen, in the last chapter, that it is incumbent on those
who maintain that all species, or even all specific characters, must be
due to natural selection, to tell us what they mean by a species, or by
characters as specific. If I am told to believe that the definite
quality A is a necessary attribute of B, and yet that B is "not a
distinct entity," but an undefinable abstraction, I can only marvel that
any one should expect me to be so simple. But, without recurring to this
point, the use of insisting on the facts above stated is, in the second
place, that otherwise I cannot suppose any general reader could believe
them in view of what is to follow. For he cannot but feel that the cost
of believing them is to render inexplicable the mental processes of
those naturalists who, in the face of such facts, have deduced the
following conclusions.
The school of naturalists against which I am contending maintains, as a
generalization deduced from the theory of natural selection, that all
species, or even all specific characters, must necessarily owe their
origin to the principle of utility. Yet this same school does not
maintain any such generalization, either with regard to varietal
characters on the one hand, or to generic characters on the other. On
the contrary, Professor Huxley, Mr. Wallace, and all other naturalists
who agree with them in refusing to entertain so much as the abstract
possibility of any cause other than natural selection having been
productive of species, fully accept the fact of other causes having been
largely concerned in the production of varieties, genera, families, and
all higher groups, or of the characters severally distinctive of each.
Indeed, Mr. Wallace does not question what appears to me the extravagant
estimate of Professor Cope, that the non-adaptive characters distinctive
of those higher groups are fully equal, in point of numbers, to the
adaptive. But, surely, if the theory of evolution by natural selection
is, as we all agree, a true theory of the origin of species, it must
likewise be a true theory of the origin of genera; and if it be supposed
essential to the integrity of the theory in its former aspect that all
specific characters should be held to be useful, I fail to see how, in
regard to its latter aspect, we are so readily to surrender the
necessary usefulness of all generic characters. And exactly the same
remark applies to the case of constant "varieties," where again the
doctrine of utility as universal is not maintained. Yet, according to
the general theory of evolution, constant varieties are what Darwin
termed "incipient species," while species are what may be termed
"incipient genera." Therefore, if the doctrine of utility as universal
be conceded to fail in the case of varieties on the one hand and of
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