Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
Until it can be shown that species are something more than merely
arbitrary divisions, due to the disappearance of intermediate varietal
links; that in some way or another they _are_ "definite entities," which
admit of being delineated by the application of some uniform or general
principles of definition; that, in short, species have only then been
classified as such when it has been shown that the origin of each has
been due to the operation of causes which have not been concerned in the
production of varieties;--until these things are shown, it clearly
remains a gratuitous dogma to maintain that forms which have been called
species differ from forms which have been called varieties in the
important respect, that they (let alone each of all their distinctive
characters) must necessarily have been due to the principle of utility.
Yet, as we have seen, even Mr. Wallace allows that a species is "not a
distinct entity," but "an assemblage of individuals which have become
somewhat modified in structure, form, and constitution"; while estimates
of the kinds and degrees of modification which are to be taken as of
specific value are conceded to be undefinable, fluctuating, and in not a
few cases almost ludicrously divergent.
Perhaps one cannot more forcibly present the rational value of this
position than by noting the following consequences of it. Mr. Gulick
writes me that while studying the land-shells of the Sandwich Islands,
and finding there a rich profusion of unique varieties, in cases where
the intermediate varieties were rare he could himself have created a
number of species by simply throwing these intermediate varieties into
his fire. Now it follows from the dogma which we are considering, that,
by so doing, not only would he have created new species, but at the same
time he would have proved them due to natural selection, and endowed the
diagnostic characters of each with a "necessarily" adaptive meaning,
which previously it was not necessary that they should present. Before
his destruction of these intermediate varieties, he need have felt
himself under no obligation to assume that any given character at either
end of the series was of utilitarian significance: but, after his
destruction of the intermediate forms, he could no longer entertain any
question upon the matter, under pain of being denounced as a Darwinian
heretic.
Public-domain text, read in full here on John Shaqi.
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