Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
Next without reference to Darwin's authority, let us see for ourselves
where the inconsistency really lies. To allow that generic characters
may be useless, while denying that specific characters can ever be so
(unless correlated with others that are useful), involves an appeal to
the argument from ignorance touching the ancestral habits,
life-conditions, &c., of a parent species now extinct. Well, even upon
this assumption of utility as obsolete, there remains to be explained
the "stability" of useless characters now distinctive of genera,
families, orders, and the rest. We know that specific characters which
have owed their origin to utility and have afterwards ceased to present
utility, degenerate, become variable, inconstant, "rudimentary," and
finally disappear. Why, then, should these things not happen with regard
to useless generic distinctions? Still more, why should they not happen
with regard to family, ordinal, and class distinctions? On the lines
against which I am arguing it would appear impossible that any answer to
this question can be suggested. For what explanation can be given of the
contrast thus presented between the obsolescence of specific characters
where previous utility is demonstrable, and the permanence of higher
characters whose previous utility is assumed? As we have already seen,
Mr. Wallace himself employs this consideration of permanence and
constancy against the view that any cause other than natural selection
can have been concerned in the origin and maintenance of _specific_
characters. But he does not seem to see that the consideration cuts two
ways--and much more forcibly against his views than in favour of them.
For while, as already shown in the chapter before last, it is
sufficiently easy to dispose of the consideration as Wallace uses it (by
simply pointing out with Darwin that any causes other than natural
selection which may have been concerned in the genesis of _specific_
characters, must, if equally uniform in their operation, equally give
rise to permanence and constancy in their results); on the other hand,
it becomes impossible to explain the stability of useless _generic_
characters, if, as Wallace's use of the argument requires, natural
selection is the only possible cause of stability. The argument is one
that cannot be played with fast and loose. Either utility is the sole
condition to the stability of _any_ diagnostic character (in which case
it is not open to Mr. Wallace to assume that all _generic_ or higher
characters which are now useless have owed their origin to a past
utility); or else utility is not the sole condition to stability (in
which case his use of the present argument in relation to _specific_
characters collapses). We have seen, indeed, in the chapter before last,
that his use of the argument collapses anyhow, or quite irrespective of
his inconsistent attitude towards generic characters, with which we were
not then concerned. But the point now is that, as a mere matter of
Public-domain text, read in full here on John Shaqi.
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