Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
if evenly distributed among all the species, need not be an adaptive
character; whereas, if any one of the species presents it in a slightly
different form, the character must be, on this account, necessarily
adaptive. In other words, the uniformity with which a generic character
occurs among the species of the genus is taken to remove that character
from the necessarily useful class, while the absence of such uniformity
is taken as proof that the character must be placed within the
necessarily useful class. Which is surely no less a _reductio ad
absurdum_ with regard to the generic character than the one just
presented with regard to its variants as specific characters. And, of
course, this twofold absurdity is presented in all cases where a generic
character is unequally distributed among the constituent species of a
genus.
[Illustration: Fig. 4.--Lower Teeth of Orang (after Tomes).]
But here is an illustration of another class of cases. Mr. Tomes has
shown that the molar teeth of the Orang present an extraordinary and
altogether superfluous amount of attachment in their sockets--the fangs
being not only exceedingly long, and therefore deeply buried in the
jaw-bone, but also curving round one another, so as still further to
strengthen the whole[133]. In the allied genera of anthropoid apes there
is no such abnormal amount of attachment. Now, the question is, of what
conceivable use can it _ever_ have been, either to the existing genus,
or to its parent species, that such an abnormal amount of attachment
should obtain? It certainly is not required to prevent dislocation of
the teeth, seeing that in all allied genera, and even in man himself,
the amount of attachment is already so great that teeth will break
before they can be drawn by anything short of a dentist's forceps.
Therefore I conclude that this peculiarity in the dentition of the genus
must have arisen in its parent species by way of what Darwin calls a
"fluctuating variation," without utilitarian significance. And I adduce
it in the present connexion because the peculiarity is one which is
equally unamenable to a utilitarian explanation, whether it happens to
occur as a generic or a specific character.
[133] _A Manual of Dental Anatomy_, p. 455.
Numberless similar cases might be quoted; but probably enough has now
been said to prove the inconsistency of the distinction which our
opponents draw between specific and all higher characters in respect of
utility. In point of fact, a very little thought is enough to show that
no such distinction admits of being drawn; and, therefore, that any one
who maintains the doctrine of utility as universal in the case of
specific characters, must in consistency hold to the same doctrine in
the case of generic and all higher characters. And the fact that our
opponents are unable to do this becomes a virtual confession on their
part of the futility of the generalization which they have
propounded[134].
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