Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
4. The doctrine in both its forms is deduced from a definition of the
theory of natural selection as a theory, and the sole theory, of the
origin of _species_; but, as Professor Huxley has already shown, it does
not really follow, even from this definition, that all specific
_characters_ must be "necessarily useful." Hence the two forms of the
doctrine, although coincident with regard to species, are at variance
with one another in respect of specific characters. Thus far, of course,
I agree with Professor Huxley; but if I have been successful in showing
that the above definition of the theory of natural selection is
logically fallacious, it follows that the doctrine in both its forms is
radically erroneous. The theory of natural selection is not, accurately
speaking, a theory of the origin of species: it is a theory of the
origin and cumulative development of adaptations, to whatever order of
taxonomic division these may happen to belong. Thus the premisses of the
deduction which we are considering collapse: the principle of utility is
shown not to have any other or further reference to species, or to
specific characters, than it has to fixed varieties, genera, families,
&c., or to the characters severally distinctive of each.
5. But, quitting all such antecedent considerations, we next proceeded
to examine the doctrine _a posteriori_, taking the arguments which have
been advanced in favour of the doctrine, other than those which rest
upon the fallacious definition. These arguments, as presented by Mr.
Wallace, are two in number.
First, it is represented that natural selection must occupy the whole
field, because no other principle of change can be allowed to operate in
the presence of natural selection. Now I fully agree that this statement
holds as regards any principle of change which is deleterious, but I
cannot agree that it does so as regards any such principle which is
merely neutral. No reason has ever been shown why natural selection
should interfere with "indifferent" characters--to adopt Professor
Huxley's term--supposing such to have been produced by any of the
agencies which we shall presently have to name. Therefore this
argument--or rather assertion--goes for nothing.
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