Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
(_c_) Altogether apart from these considerations, there remains a much
more important one. For the objection that fixed--or "stable"--climatic
varieties differ from true species in not being subject to heredity,
raises the question--What are we to understand by a "species"? This
question, which was thus far purposely left in abeyance, had now to be
dealt with seriously. For it would clearly be irrational in our
opponents to make this highly important generalization with regard to
species and specific characters, unless they are prepared to tell us
what they mean by species, and therefore by characters as specific. In
as far as there is any ambiguity on this point it makes entirely for
our side in the debate, because even any small degree of uncertainty
with regard to it would render the generalization in question
proportionally unsound. Yet it is notorious that no word in existence is
more vague, or more impossible to define, than the word "species." The
very same men who at one time pronounce their great generalization with
regard to species, at another time asseverate that "a species is not a
definite entity," but a merely abstract term, serving to denote this
that and the other organic type, which this that and the other
systematist regards as deserving such a title. Moreover it is
acknowledged that systematists differ among themselves to a wide extent
as to the kinds and degrees of peculiarity which entitle a given form to
a specific rank. Even in the same department of systematic work much
depends on merely individual taste, while in different departments
widely different standards of delimination are in vogue. Hence, our
_reductio ad absurdum_ consists in this--that whether a given form is to
be regarded as necessarily due to natural selection, and whether all its
distinctive characters are to be regarded as necessarily utilitarian
characters, will often depend on whether it has been described by
naturalist A or by naturalist B. There is no one criterion--there is not
even any one set of criteria--agreed upon by naturalists for the
construction of specific types. In particular, as regards the principle
of heredity, it is not known of one named species in twenty--probably
not in a hundred--whether its diagnostic characters are hereditary
characters; while, on the other hand, even in cases where experiment has
proved "constant varieties" to be hereditary--and even also
cross-sterile with allied varieties--it is only some three or four
living botanists who for these reasons advocate the elevation of such
varieties to the rank of species. In short, as we are not engaged on any
abstract question touching the principles on which species ought to have
been constituted by their makers, but upon the actual manner in which
they have been, the criterion of heredity must needs be disregarded in
the present discussion, as it has been in the work of systematists. And
the result of this is, that any objection to our introducing the facts
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