Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
"To account for the diminution of organs or structures no longer of
use, apart from any inherited effects of disuse, Mr. Romanes has
invoked the Cessation of Selection; and Mr. Francis Galton has, in
another connexion, summarized the effects of this cessation of
selection in the convenient phrase 'Regression to Mediocrity.' This
is the Panmixia of Professor Weismann and his followers; but the
phrase regression to mediocrity through the cessation of selection
appears to me preferable. It is clear that so long as any organ or
structure is subject to natural selection through elimination, it
is, if not actually undergoing improvement, kept at a high standard
of efficiency through the elimination of all those individuals in
which the organ in question falls below the required standard. But
if, from change in the environment or any other cause, the
character in question ceases to be subject to selection,
elimination no longer takes place, and the high standard will no
longer be maintained. There will be reversion to mediocrity. The
probable amount of this reversion is at present a matter under
discussion[143]."
[143] _Presidential Address to the Bristol Naturalists' Society_,
1891.
So far, then, Professor Lloyd Morgan is in complete agreement with
previous writers upon the subject. He does not doubt that the cessation
of selection must always be a cause of degeneration: the only question
is as to the _potency_ of this cause, or the _amount_ of degeneration
which it is capable of effecting.
Taking, first, the case of bulk or size of an organ, as distinguished
from its organization or complexity, we have seen that Weismann
represents the cessation of selection--even if working quite alone, or
without any assistance from the reversal of selection--to be capable of
reducing a fully developed organ to the state of a rudiment, or even, if
we take his most recent view, of abolishing the organ _in toto_.
Professor Lloyd Morgan, on the other hand, does not think that the
cessation of selection alone can cause reduction further than the level
of "mediocrity" in the first generations--or, which is much the same
thing, further than the difference between the "birth-mean" and the
"selection-mean" of the first generations. This amount of reduction he
puts at 5 per cent., as "a very liberal estimate."
Public-domain text, read in full here on John Shaqi.
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