Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
As far as I am aware, the only occasion on which Professor Huxley has
alluded to the subject in question, is in his obituary notice of Darwin
in the _Proceedings of the Royal Society_, Vol. XLIV, No. 269, p. xviii.
The allusion is to my paper on _Physiological Selection_, in the
_Journal of the Linnæan Society_, Zool. Vol. XIX, pp. 337-411. But it
will be observed that the criticism has no reference to the theory which
it is the object of that paper to set forth. It refers only to my
definition of the theory of natural selection as primarily a theory of
the origin, or cumulative development, of adaptations. This criticism,
together with my answer thereto at the time, is conveyed in the
following words.
"Every variety which is selected into a species is favoured and
preserved in consequence of being, in some one or more respects,
better adapted to its surroundings than its rivals. In other words,
every species which exists, exists in virtue of adaptation, and
whatever accounts for that adaptation accounts for the existence of
the species. To say that Darwin has put forward a theory of the
adaptation of species, but not of their origin, is therefore to
misunderstand the first principles of the theory. For, as has been
pointed out, it is a necessary consequence of the theory of
selection that every species must have some one or more structural
or functional peculiarities, in virtue of the advantage conferred
by which it has fought through the crowd of its competitors, and
achieved a certain duration. In this sense, it is true that every
species has been 'originated' by selection."
Now, in the first place, I have nowhere said that "Darwin has put
forward a theory of the adaptation of species, but not of their
origin." I said, and continue to say, that he has put forward a
theory of _adaptations in general_, and that where such adaptations
appertain to species only (i.e. are peculiar to particular
species), the theory becomes "_also_ a theory of the origin of the
species which present them." The only possible misunderstanding,
therefore, which can here be alleged against me is, that I fail to
perceive it as a "necessary consequence of the theory of selection
that _every_ species _must_ have some one or more structural or
functional _peculiarities_" of an adaptive or utilitarian kind.
Now, if this is a misunderstanding, I must confess to not having
had it removed by Mr. Huxley's exposition.
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