Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
That Mr. Wallace himself should be biassed in this matter might,
perhaps, be expected. After rendering the following very unequivocal
passage from the _Origin of Species_ (p. 72)--"There can be little doubt
that the tendency to vary in the same manner has often been so strong,
_that all individuals of the same species have been similarly modified
without the aid of any form of selection_"--Mr. Wallace says, "But no
proof whatever is offered of this statement, and it is so entirely
opposed to all we know of the facts of variation as given by Darwin
himself, that the important word 'all' is probably an oversight." But,
if Mr. Wallace had read the very next sentence he would have seen that
here the important word "all" could not _possibly_ have been "an
oversight." For the passage continues,--"Or only a third, fifth, or
tenth part of the individuals may have been thus affected, of which fact
several instances could be given. Thus Graba estimates that about
one-fifth of the guillemots in the Faroe Islands consist of a variety so
well marked, that it was formerly ranked as a distinct species under the
name of Uria lacrymans." And even if this passage had not been thus
specially concerned with the question of the _proportion_ in which
"_individuals of the same species have been similarly modified without
the aid of any form of selection_" the oversight with respect to "the
important word 'all'" would still have remained an oversight of a
recurrent character, as the following additional quotations from other
parts of Darwin's writings may perhaps render apparent.
"There must be some efficient cause for each slight individual
difference, as well as for more strongly marked variations which
occasionally arise; and if the unknown cause were to act
persistently, it is almost certain that _all_ the individuals of
the _species_ would be similarly modified[168]."
"The acquisition of a useless part can hardly be said to raise an
organism in the natural scale.... We are so ignorant of the
exciting cause of the above specified modifications; but if the
unknown cause were to act almost uniformly for a length of time, we
may infer that the result would be almost uniform; and in this case
_all_ the individuals of the _species_ would be modified in the
same manner[169]."
[168] _Origin of Species_, p. 171.
[169] _Ibid._ p. 175.
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