Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
But, in the second place, there is a wide difference between the
certainty of this fact and that of the theory based upon it. The certain
fact is, that a great distinction in respect of heritability is
observable between congenital and acquired characters. The theory, as
formulated by Weismann, is that the distinction is not only great but
absolute, or, in other words, that in no case and in no degree can any
acquired character be ever inherited. This hypothesis, it will be
observed, goes far beyond the observed fact, for it is obviously
possible that, notwithstanding this great difference in regard to
heritability between congenital and acquired characters, the latter may
nevertheless, sometimes and in some degree, be inherited, however much
difficulty we may experience in observing these lesser phenomena in
presence of the greater. The Weismannian hypothesis of _absolute_
continuity is one thing, while the observed fact of at least a _high
relative degree_ of continuity is quite another thing. And it is
necessary to be emphatic on this point, since some of the reviewers of
my _Examination of Weismannism_ confound these two things. Being
apparently under the impression that it was reserved for Weismann to
perceive the fact of there being a great difference between the
heritability of congenital and acquired characters, they deem it
inconsistent in me to acknowledge this fact while at the same time
questioning the hypothetical basis of his fundamental postulate touching
the absolute continuity of germ-plasm. It is one merit of Galton's
theory, as against Weismann's, that it does not dogmatically exclude the
possible interruption of continuity on some occasions and in some
degree. Herein, indeed, would seem to lie the central core of the whole
question in dispute. For it is certain and has long been known that
individually acquired characters are at all events much less heritable
than are long-inherited or congenital ones. But Lamarckian theory
supposes that congenital characters were in some cases originally
acquired, and that what are now blastogenetic characters were in some
cases at first somatogenetic and have become blastogenetic only in
virtue of sufficiently long inheritance. Since Darwin's time, however,
evolutionists (even of the so-called Lamarckian type) have supposed that
natural selection greatly assists this process of determining which
somatogenetic characters shall become congenital or blastogenetic. Hence
all schools of evolutionists are, and have long been, agreed in
regarding the continuity principle as true in the main. No evolutionist
would at any time have propounded the view that one generation depends
for _all_ its characters on those acquired by its _immediate_ ancestors,
for this would merely be to unsay the theory of Evolution itself, as
well as to deny the patent facts of heredity as shown, for example, in
atavism. At most only some fraction of a _per cent._ could be supposed
to do so.
Public-domain text, read in full here on John Shaqi.
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