Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
in the vegetable kingdom) that we find the most
unequivocal evidence of its action; while, on the other hand, it is just
where it can be seen to have the least opportunity of asserting itself
(viz. among the higher animals) that we find it most associated with,
and therefore assisted by, other forms of homogamy, i. e. not only
geographical isolation, but also by sexual preference in pairing, and
the several other forms of homogamy, which Mr. Gulick has shown to arise
in different places as the result of intelligence.
_Evidence from Special Cases._
Hitherto I have been considering, from the most general point of view,
the most widespread facts and broadest principles which serve to
substantiate the theory of physiological selection. I now pass to the
consideration of one of those special cases in which the theory appears
to have been successfully applied.
Professor Le Conte has adduced the fossil snails of Steinheim as serving
to corroborate the theory of physiological selection[26].
[26] _Evolution and its Relations to Religious Thought_, &c. pp.
236-7.
The facts are these. The snail population of this lake remain for a long
time uniform and unchanged. Then a small percentage of individuals
suddenly began to vary as regards the form of their shells, and this in
two or three directions at the same time, each affected individual,
however, only presenting one of the variations. But after all these
variations had begun to affect a proportionally large number of
individuals, some individuals occur in which two or more of the
variations are blended together, evidently, as Weismann says, by
intercrossing of the varieties so blended. Later still, both the
separate varieties and their blended progeny became more and more
numerous, and eventually a single blended type, comprising in itself all
the initial varieties, supplanted the parent form. Then another long
period of stability ensued until another eruption of new variations took
place; and these variations, after having affected a greater and greater
number of individuals, eventually blended together by intercrossing and
supplanted their parent form. So the process went on, comparatively
short periods of variation alternating with comparatively long periods
of stability, the variations, moreover, always occurring suddenly in
crops, then multiplying, blending together, and in their finally blended
type eventually supplanting their parent form.
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