Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
Here we have a curious example of one writer supporting the statements
of another, while appearing to be under the impression that he is
controverting those statements. Both Professor Herdman's imaginary case,
and its realization in Livingstone's account, go to show "the tendency
of the members of a variety to breed with one another." This is what I
have called "psychological selection," and, far from "ignoring" it, I
have always laid stress upon it as an obviously important form of
isolation or _prevention_ of free intercrossing. But it is a form of
isolation which can only occur in the higher animals, and, therefore,
the whole of Professor Herdman's criticism is merely a restatement of my
own views as already published in the paper which he is criticizing.
For all that his argument goes to prove is, first, the necessity for
_some_ form of isolation if the overwhelming effects of intercrossing
are to be obviated; and, secondly, the manifest consequence that where
the psychological form is unavailable (as in many of the lower animals
and in all plants), some other form must be present if divergent
evolution is taking place on a common area.
* * * * *
Seeing that so much misunderstanding has been shown with reference to my
views on "the swamping effects of intercrossing," and seeing also that
this misunderstanding extends quite as much to Mr. Gulick's views as to
my own, I will here supply brief extracts from both our original papers,
for the double purpose of showing our complete agreement, and of leaving
it to be judged whether we can fairly be held responsible for the
misunderstanding in question. After having supplied these quotations, I
will conclude this historical sketch by considering what Mr. Wallace has
said in reply to the views therein presented. I will transcribe but a
single passage from our papers, beginning with my own.
Any theory of the origin of species in the way of descent must be
prepared with an answer to the question, Why have species
_multiplied_? How is it that, in the course of evolution, species
have not simply become transmuted in linear series instead of
ramifying into branches? This question Mr. Darwin seeks to answer
"from the simple circumstance that the more diversified the
descendants from any one species becomes in structure,
constitution, and habits, by so much will they be better enabled to
seize on many and widely diversified places in the economy of
nature, and so be enabled to increase in numbers." And he proceeds
to illustrate this principle by means of a diagram, showing the
hypothetical divergence of character undergone by the descendants
of seven species. Thus, he attributes divergence of character
exclusively to the influence of natural selection.
Public-domain text, read in full here on John Shaqi.
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