Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
That perfectly free intercrossing, between any or all of the
individuals of a given group of animals, is, so long as the
characters of the parents are blended in the offspring, fatal to
divergence of character, is undeniable. Through the elimination of
less favourable variations, the swiftness, strength, and cunning of
a race may be gradually improved. But no form of elimination can
possibly differentiate the group into swift, strong, and cunning
varieties, distinct from each other, so long as all three varieties
freely interbreed, and the characters of the parents blend in the
offspring. Elimination may and does give rise to progress in any
given group, _as a group_; it does not and cannot give rise to
differentiation and divergence, so long as interbreeding with
consequent interblending of characters be freely permitted. Whence
it inevitably follows, as a matter of simple logic, that where
divergence has occurred, intercrossing and interbreeding must in
some way have been lessened or prevented. Thus a new factor is
introduced, that of _isolation_ or _segregation_. And there is no
questioning the fact that it is of great importance. Its
importance, indeed, can only be denied by denying the swamping
effects of intercrossing, and such denial implies the tacit
assumption that interbreeding and interblending are held in check
by some form of segregation. The isolation explicitly denied is
implicitly assumed[51].
[51] _Animal Life and Intelligence_, pp. 98, 99 (1890-1891).
Similarly, and still more recently, Professor Le Conte writes:--
It is evident, then, as Romanes claims, that natural selection
alone tends to _monotypic_ evolution. Isolation of some sort seems
necessary to _polytypic_ evolution. The tree of evolution under the
influence of natural selection alone grows palm-like from its
terminal bud. Isolation was necessary to the starting of lateral
buds, and thus for the profuse ramification which is its most
conspicuous character[52].
[52] _The Factors of Evolution_ (1891).
In order to complete this historical review, it only remains to consider
Mr. Wallace's utterances upon the subject.
Public-domain text, read in full here on John Shaqi.
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