Darwin, and After Darwin, Volumes 1 and 3: An Exposition of the Darwinian Theory and a Discussion of Post-Darwinian QuestionsRomanes, George John
Science
Darwin, and After Darwin, Volumes 1 and 3: An Exposition of the Darwinian Theory and a Discussion of Post-Darwinian Questions
Romanes, George John
Evolution (Biology); Heredity
The above is a brief condensation of the already condensed statement
which Darwin has given of the imperfection of the geological record; but
I think it is enough to show, in a general way, how precarious must be
the nature of any objections to the theory of evolution which are
founded merely upon the silence of palæontology in cases where, if the
record were anything like complete, we should be entitled to expect from
it some positive information. But, as we have seen in the text,
imperfect though the record be, in as far as it furnishes positive
information at all, this is well-nigh uniformly in favour of the theory;
and therefore, even on grounds of palæontology alone, it appears to me
that Darwin is much too liberal where he concludes his discussion by
saying,--"Those who believe that the geological record is in any degree
perfect, will undoubtedly at once reject the theory." If in any measure
reasonable, such persons ought rather to examine their title to such a
belief; and even if they disregard the consensus of testimony which is
yielded by all the biological sciences to the theory of evolution, they
ought at least to hold their judgment in suspense until they shall have
not only set against the apparently negative testimony which is yielded
by geology its unquestionably positive testimony, but also well
considered the causes which may--or rather must--have so gravely
impaired the geological record.
However, be this as it may, I will now pass on to consider the
difficulties and objections which have been brought against the theory
on grounds of palæontology.
These may be classified under four heads. First, the absence of varietal
links between allied species; second, the sudden appearance of whole
groups of species--not only as genera and families, but even sometimes
as orders and classes--without any forms leading up to them; third, the
occurrence of highly organized types at much lower levels of geological
strata than an evolutionist would antecedently expect; and, fourth, the
absence of fossils of any kind lower down than the Cambrian strata.
Now all these objections depend on estimates of the imperfection of the
geological record much lower than that which is formed by Darwin.
Therefore I have arranged the objections in their order of difficulty in
this respect, or in the order that requires successively increasing
estimates of the imperfection of the record, if they are to be
successively answered.
I think that the first of them has been already answered in the text, by
showing that even a very moderate estimate of the imperfection of the
record is enough to explain why intermediate _varieties_, connecting
allied _species_, are but comparatively seldom met with. Moreover it was
shown that in some cases, where shells are concerned, remarkably
well-connected series of such varieties have been met with. And the same
applies to species and genera in certain other cases, as in the equine
family.
Public-domain text, read in full here on John Shaqi.
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