Darwin, and After Darwin, Volumes 1 and 3: An Exposition of the Darwinian Theory and a Discussion of Post-Darwinian QuestionsRomanes, George John
Science
Darwin, and After Darwin, Volumes 1 and 3: An Exposition of the Darwinian Theory and a Discussion of Post-Darwinian Questions
Romanes, George John
Evolution (Biology); Heredity
The case most frequently insisted on by palæontologists of the
apparently sudden appearance of a whole group of species, is that
of the teleostean fishes, low down, according to Agassiz, in the
Chalk period. This group includes the large majority of existing
species. But certain Jurassic and Triassic forms are now commonly
admitted to be teleostean; and even some palæozoic forms have been
thus classed by one high authority. If the teleosteans had really
appeared suddenly in the northern hemisphere, the fact would have
been highly remarkable; but it would not have formed an insuperable
difficulty, unless it could likewise have been shown that at the
same period the species were suddenly and simultaneously developed
in other quarters of the world. It is almost superfluous to remark
that hardly any fossil fish are known from south of the equator;
and by running through Pictet's Palæontology it will be seen that
very few species are known from several formations in Europe. Some
few families of fish now have a confined range; the teleostean fish
might formerly have had a similarly confined range, and after
having been largely developed in some one sea, might have spread
widely. Nor have we any right to suppose that the seas of the world
have always been so freely open from south to north as they are at
present. Even at this day, if the Malay Archipelago were converted
into land, the tropical parts of the Indian Ocean would form a
large and perfectly enclosed basin, in which any great group of
marine animals might be multiplied; and here they would remain
confined, until some of the species became adapted to a cooler
climate, and were enabled to double the southern capes of Africa or
Australia, and thus reach other and distant seas.
From these considerations, from our ignorance of the geology of
other countries beyond the confines of Europe and the United
States; and from the revolution in our palæontological knowledge
effected by the discoveries of the last dozen years, it seems to me
to be about as rash to dogmatize on the succession of organic
forms throughout the world, as it would be for a naturalist to
land for five minutes on some one barren point in Australia, and
then to discuss the number and range of its productions[53].
[53] _Origin of Species_, 282-5.
In view of all the foregoing facts and considerations, it appears to me
that the second difficulty on our list is completely answered. Indeed,
even on a moderate estimate of the imperfection of the geological
record, the wonder would have been if many cases had _not_ occurred
where groups of species present the fictitious appearance of having been
suddenly and simultaneously created in the particular formations where
their remains now happen to be observable.
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