It must, therefore, be admitted that one outcome of the progress of embryological and palaeontological research for the last 50 years is negative. The recapitulation theory originated as a deduction from the evolution theory and as a deduction it still remains. Let us before leaving the subject apply another test. If the evolution theory and the recapitulation theory are both true, how is it that living birds are not only without teeth but have no rudiments of teeth at any stage of their existence? How is it that the missing digits in birds and mammals, the missing or reduced limb of snakes and whales, the reduced mandibulo-hyoid cleft of elasmobranch fishes are not present or relatively more highly developed in the embryo than in the adult? How is it that when a marked variation, such as an extra digit, or a reduced limb, or an extra segment, makes its appearance, it is not confined to the adult but can be seen all through the development? All the clear evidence we can get tends to show that marked variations, whether of reduction or increase, of organs are manifest during the whole of the development of the organ and do not merely affect the adult. And on reflection we see that it could hardly be otherwise. All such evidence is distinctly at variance with the theory of recapitulation, at least as applied to embryos. In the case of larvae of course the case will be different, for in them the organs are functional, and reduction in the adult will not be accompanied by reduction in the larva unless a change in the conditions of life of the larva enables it to occur. If after 50 years of research and close examination of the facts of embryology the recapitulation theory is still without satisfactory proof, it seems desirable to take a wider sweep and to inquire whether the facts of embryology cannot be included in a larger category.