Darwin and Modern ScienceSeward, A. C. (Albert Charles)
Philosophy
Darwin and Modern Science
Seward, A. C. (Albert Charles)
Darwin, Charles, 1809-1882; Evolution (Biology); Science -- History
Abnormal developments of all kinds in stems, leaves, and flowers, may
be produced by parasites, insects, or fungi. They may also be induced
by injury, as Blaringhem (Blaringhem, "Mutation et traumatismes", Paris,
1907.) has more particularly demonstrated, which, by cutting away the
leading shoots of branches in an early stage of development, caused
fasciation, torsion, anomalous flowers, etc. The experiments of
Blaringhem point to the probability that disturbances in the conditions
of food-supply consequent on injury are the cause of the production of
monstrosities. This is certainly the case in my experiments with species
of Sempervivum (Klebs, "Kunstliche Metamorphosen", Stuttgart, 1906.);
individuals, which at first formed normal flowers, produced a great
variety of abnormalities as the result of changes in nutrition, we
may call to mind the fact that the formation of inflorescences occurs
normally when a vigorous production of organic compounds, such as
starch, sugar, etc. follows a diminution in the supply of mineral
salts. On the other hand, the development of inflorescences is entirely
suppressed if, at a suitable moment before the actual foundations have
been laid, water and mineral salts are supplied to the roots. If, during
the week when the inflorescence has just been laid down and is growing
very slowly, the supply of water and salts is increased, the internal
conditions of the cells are essentially changed. At a later stage, after
the elongation of the inflorescence, rosettes of leaves are produced
instead of flowers, and structures intermediate between the two kinds of
organs; a number of peculiar plant-forms are thus obtained (Cf. Lotsy,
"Vorlesungen uber Deszendenztheorien", Vol. II. pl. 3, Jena, 1908.)
Abnormalities in the greatest variety are produced in flowers by varying
the time at which the stimulus is applied, and by the cooperation
of other factors such as temperature, darkness, etc. In number and
arrangement the several floral members vary within wide limits;
sepals, petals, stamens, and carpels are altered in form and colour, a
transformation of stamens to carpels and from carpels to stamens occurs
in varying degrees. The majority of the deviations observed had not
previously been seen either under natural conditions or in cultivation;
they were first brought to light through the influence of external
factors.
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