the same time developed in the insect the instinct to press the pollen into the pistil. In this domain, then, the theory of selection finds nothing but corroboration, and it would be impossible to substitute for it any other explanation, which, now that the facts are so well known, could be regarded as a serious rival to it. That selection is a factor, and a very powerful factor in the evolution of organisms, can no longer be doubted. Even although we cannot bring forward formal proofs of it IN DETAIL, cannot calculate definitely the size of the variations which present themselves, and their selection-value, cannot, in short, reduce the whole process to a mathematical formula, yet we must assume selection, because it is the only possible explanation applicable to whole classes of phenomena, and because, on the other hand, it is made up of factors which we know can be proved actually to exist, and which, IF they exist, must of logical necessity cooperate in the manner required by the theory. WE MUST ACCEPT IT BECAUSE THE PHENOMENA OF EVOLUTION AND ADAPTATION MUST HAVE A NATURAL BASIS, AND BECAUSE IT IS THE ONLY POSSIBLE EXPLANATION OF THEM. (This has been discussed in many of my earlier works. See for instance "The All-Sufficiency of Natural Selection, a reply to Herbert Spencer", London, 1893.) Many people are willing to admit that selection explains adaptations, but they maintain that only a part of the phenomena are thus explained, because everything does not depend upon adaptation. They regard adaptation as, so to speak, a special effort on the part of Nature, which she keeps in readiness to meet particularly difficult claims of the external world on organisms. But if we look at the matter more carefully we shall find that adaptations are by no means exceptional, but that they are present everywhere in such enormous numbers, that it would be difficult in regard to any structure whatever, to prove that adaptation had NOT played a part in its evolution. How often has the senseless objection been urged against selection that it can create nothing, it can only reject. It is true that it cannot create either the living substance or the variations of it; both must be given. But in rejecting one thing it preserves another, intensifies it, combines it, and in this way CREATES what is new. EVERYTHING in organisms depends on adaptation; that is to say, everything must be admitted through the narrow door of selection, otherwise it can take no part in the building up of the whole. But, it is asked, what of the direct effect of external conditions, temperature, nutrition, climate and the like? Undoubtedly these can give rise to variations, but they too must pass through the door of selection, and if they cannot do this they are rejected, eliminated from the constitution of the species.