This is safe and to a certain degree judicious, no doubt, as respects
published species. Once admitted, they may stand until they are put down by
evidence, direct or circumstantial. Doubtless a species may rightfully be
condemned on good circumstantial evidence. But what course does De Candolle
pursue in the case--of every-day occurrence to most working botanists,
having to elaborate collections from countries not so well explored as
Europe--when the forms in question, or one of the two, are as yet unnamed?
Does he introduce as a new species every form which he cannot connect by
ocular proof with a near relative, from which it differs only in
particulars which he sees are inconstant in better known species of the
same group? We suppose not. But, if he does, little improvement for the
future upon the state of things revealed in the following quotation can be
expected:
"In the actual state of our knowledge, after having seen nearly all the
original specimens, and in some species as many as two hundred
representatives from different localities, I estimate that, out of the
three hundred species of Cupuliferae which will be enumerated in the
Prodromus, two-thirds at least are provisional species. In general, when we
consider what a multitude of species were described from a single specimen,
or from the forms of a single locality, of a single country, or are badly
described, it is difficult to believe that above one-third of the actual
species in botanical works will remain unchanged."
Such being the results of the want of adequate knowledge, how is it likely
to be when our knowledge is largely increased? The judgment of so practised
a botanist as De Candolle is important in this regard, and it accords with
that of other botanists of equal experience.
"They are mistaken," he pointedly asserts, "who repeat that the greater
part of our species are clearly limited, and that the doubtful species are
in a feeble minority. This seemed to be true, so long as a genus was
imperfectly known, and its species were founded upon few specimens, that is
to say, were provisional. Just as we come to know them better, intermediate
forms flow in, and doubts as to specific limits augment."
Public-domain text, read in full here on John Shaqi.
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