Before the publication of Darwin's first volume, M. Alphonse de Candolle
had summed up the result of his studies in this regard, in the final
chapter of his classical "Geographie Botanique Raisonnee," in the
conclusion, that existing vegetation must be regarded as the continuation,
through many geological and geographical changes, of the anterior
vegetations of the world; and that, consequently, the present distribution
of species is explicable only in the light of their geological history. He
surmised that, notwithstanding the general stability of forms, certain
species or quasi-species might have originated through diversification
under geographical isolation. But, on the other hand, he was still disposed
to admit that even the same species might have originated independently in
two or more different regions of the world; and he declined, as unpractical
and unavailing, all attempts to apply hypotheses to the elucidation of the
origin of species. Soon after Darwin's book appeared, De Candolle had
occasion to study systematically a large and wide-spread genus-- that of
the oak. Investigating it under the new light of natural selection, he came
to the conclusion that the existing oaks are all descendants of earlier
forms, and that no clear line can be drawn between the diversification whic
h has resulted in species and that which is exhibited in races and minor
varieties.
And now, in the introductory chapter of the volume of essays before us, he
informs us that the idea which pervades them all, and in some sort connects
very diverse topics, is that of considering this principle of selection. Of
the principle itself, he remarks that it is neither a theory nor an
hypothesis, but the expression of a necessary fact; that to deny it is very
much like denying that round stones will roll downhill faster and farther
than flat ones; and that the question of the present day in natural history
is not whether there be natural selection, or even whether forms are derived
from other forms, but to comprehend how, in what proportions, and by what
means hereditary deviations take place, and in what ways an inevitable
selection takes effect upon these. In two of these essays natural selection
is directly discussed in its application to the human race; the larger one
dealing ably with the whole subject, and with results at first view
seemingly in a great degree negative, but yet showing that the supposed
"failure of natural selection in the case of man" was an unwarrantable
conclusion from too limited a view of a very complicated question. The
article abounds in acute and fertile suggestions, and its closing chapter,
"on the probable future of the human species" under the laws of selection,
is highly interesting and noteworthy. The other and shorter essay discusses
a special point, and brings out a corollary of the law of heredity which may
not have been thought of before, but which is perfectly clear as soon as it
is stated.
Public-domain text, read in full here on John Shaqi.
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