This language implies that the animal matter is in some way or other
discerned by the tentacles, and is appropriated. Formerly there was only a
presumption of this, on the general ground that such an organization could
hardly be purposeless. Yet, while such expressions were natural, if not
unavoidable, they generally were used by those familiar with the facts in a
half-serious, half-metaphorical sense. Thanks to Mr. Darwin's
investigations, they may now be used in simplicity and seriousness.
That the glands secrete the glairy liquid of the drop is evident, not only
from its nature, but from its persistence through a whole day's exposure to
a summer sun, as also from its renewal after it has been removed, dried up,
or absorbed. That they absorb as well as secrete, and that the whole
tentacle may be profoundly affected thereby, are proved by the different
effects, in kind and degree, which follow the application of different
substances. Drops of rain-water, like single momentary touches of a solid
body, produce no effect, as indeed they could be of no advantage; but a
little carbonate of ammonia in the water, or an infusion of meat, not only
causes inflection, but promptly manifests its action upon the contents of
the cells of which the tentacle is constructed. These cells are
sufficiently transparent to be viewed under the microscope without
dissection or other interference; and the change which takes place in the
fluid contents of these cells, when the gland above has been acted upon, is
often visible through a weak lens, or sometimes even by the naked eye,
although higher powers are required to discern what actually takes place.
This change, which Mr. Darwin discovered, and turns to much account in his
researches, he terms "aggregation of the protoplasm." When untouched and
quiescent, the contents appear as an homogeneous purple fluid. When the
gland is acted upon, minute purple particles appear, suspended in the now
colorless or almost colorless fluid; and this change appears first in the
cells next the gland, and then in those next beneath, traveling down the
whole length of the tentacle. When the action is slight, this appearance
does not last long; the particles of "aggregated protoplasm redissolved, the
process of redissolution traveling upward from the base of the tentacle to
the gland in a reverse direction to that of the aggregation. Whenever the
action is more prolonged or intense, as when a bit of meat or crushed fly,
or a fitting solution, is left upon the gland, the aggregation proceeds
further, so that the whole protoplasm of each cell condenses into one or
two masses, or into a single mass which will often separate into two, which
afterward reunite; indeed, they incessantly change their forms and
positions, being never at rest, although their movements are rather slow.
In appearance and movements they are very like amoebae and the white
corpuscles of the blood. Their motion, along with the streaming movement of
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