But the remaining species, S. variolaris, is the most wonderful of our
pitcher-plants in its adaptations for the capture of insects. The inflated
and mottled lid or hood overarches the ample orifice of the tubular pitcher
sufficiently to ward off the rain, but not to obstruct the free access of
flying insects. Flies, ants, and most insects, glide and fall from the
treacherous smooth throat into the deep well below, and never escape. They
are allured by a sweet secretion just within the orifice-- which was
discovered and described long ago, and the knowledge of it wellnigh
forgotten until recently. And, finally, Dr. Mellichamp, of South Carolina,
two years ago made the capital discovery that, during the height of the
season, this lure extends from the orifice down nearly to the ground, a
length of a foot or two, in the form of a honeyed line or narrow trail on
the edge of the wing-like border which is conspicuous in all these species,
although only in this one, so far as known, turned to such account. Here,
one would say, is a special adaptation to ants and such terrestrial and
creeping insects. Well, long before this sweet trail was known, it was
remarked by the late Prof. Wyman and others that the pitchers of this
species, in the savannahs of Georgia and Florida, contain far more ants
than they do of all other insects put together.
Finally, all this is essentially repeated in the peculiar Californian
pitcher-plant (Darlingtonia), a genus of the same natural family, which
captures insects in great variety, enticing them by a sweetish secretion
over the whole inside of the inflated hood and that of a curious forked
appendage, resembling a fish-tail, which overhangs the orifice. This
orifice is so concealed that it can be seen and approached only from below,
as if--the casual observer might infer--to escape visitation. But dead
insects of all kinds, and their decomposing remains, crowd the cavity and
saturate the liquid therein contained, enticed, it is said, by a peculiar
odor, as well as by the sweet lure which is at some stages so abundant as
to drip from the tips of the overhanging appendage. The principal
observations upon this pitcher-plant in its native habitat have been made
by Mrs. Austin, and only some of the earlier ones have thus far been
published by Mr. Canby. But we are assured that in this, as in the
Sarracenia variolaris, the sweet exudation extends at the proper season
from the orifice down the wing nearly to the ground, and that ants follow
this honeyed pathway to their destruction. Also, that the watery liquid in
the pitcher, which must be wholly a secretion, is much increased in
quantity after the capture of insects.
Public-domain text, read in full here on John Shaqi.
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