Under this view, moreover, waste of life and material in organic Nature
ceases to be utterly inexplicable, because it ceases to be objectless. It
is seen to be a part of the general "economy of Nature," a phrase which has
a real meaning. One good illustration of it is furnished by the pollen of
flowers. The seeming waste of this in a pine-forest is enormous. It gives
rise to the so-called "showers of sulphur," which every one has heard of.
Myriads upon myriads of pollen-grains (each an elaborate organic structure)
are wastefully dispersed by the winds to one which reaches a female flower
and fertilizes a seed. Contrast this with one of the close-fertilized
flowers of a violet, in which there are not many times more grains of
pollen produced than there are of seeds to be fertilized; or with an
orchis-flower, in which the proportion is not widely different. These
latter are certainly the more economical; but there is reason to believe
that the former arrangement is not wasteful. The plan in the violet-flower
assures the result with the greatest possible saving of material and
action; but this result, being close-fertilization or breeding in and in,
would, without much doubt, in the course of time, defeat the very object of
having seeds at all.[XIII-3] So the same plant produces other flowers also,
provided with a large surplus of pollen, and endowed (as the others are
not) with color, fragrance, and nectar, attractive to certain insects, which
are thereby induced to convey this pollen from blossom to blossom, that it
may fulfill its office. In such blossoms, and in the great majority of
flowers, the fertilization and consequent perpetuity of which are committed
to insects, the likelihood that much pollen may be left behind or lost in
the transit is sufficient reason for the apparent superfluity. So, too, the
greater economy in orchis-flowers is accounted for by the fact that the
pollen is packed in coherent masses, all attached to a common stalk, the end
of which is expanded into a sort of button, with a glutinous adhesive face
(like a bit of sticking-plaster), and this is placed exactly where the head
of a moth or butterfly will be pressed against it when it sucks nectar from
the flower, and so the pollen will be bodily conveyed from blossom to
blossom, with small chance of waste or loss. The floral world is full of
such contrivances; and while they exist the doctrine of purpose or final
cause is not likely to die out. Now, in the contrasted case, that of
pine-trees, the vast superabundance of pollen would be sheer waste if the
intention was to fertilize the seeds of the same tree, or if there were any
provision for insect-carriage; but with wide-breeding as the end, and the
wind which "bloweth where it listeth" as the means, no one is entitled to
declare that pine-pollen is in wasteful excess. The cheapness of
wind-carriage may be set against the overproduction of pollen.
Public-domain text, read in full here on John Shaqi.
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