Darwiniana : Essays — Volume 02Huxley, Thomas Henry
Science
Darwiniana : Essays — Volume 02
Huxley, Thomas Henry
Darwin, Charles, 1809-1882; Evolution (Biology)
The conclusions enunciated by Cuvier and Von Baer have been confirmed, in
principle, by all subsequent research into the structure of animals and
plants. But the effect of the adoption of these conclusions has been rather
to substitute a new metaphor for that of Bonnet than to abolish the
conception expressed by it. Instead of regarding living things as capable
of arrangement in one series like the steps of a ladder, the results of
modern investigation compel us to dispose them as if they were the twigs
and branches of a tree. The ends of the twigs represent individuals, the
smallest groups of twigs species, larger groups genera, and so on, until we
arrive at the source of all these ramifications of the main branch, which
is represented by a common plan of structure. At the present moment, it is
impossible to draw up any definition, based on broad anatomical or
developmental characters, by which any one of Cuvier's great groups shall
be separated from all the rest. On the contrary, the lower members of each
tend to converge towards the lower members of all the others. The same may
be said of the vegetable world. The apparently clear distinction between
flowering and flowerless plants has been broken down by the series of
gradations between the two exhibited by the _Lycopodiaceæ,
Rhizocarpeæ_, and _Gymnospermeæ_. The groups of _Fungi_,
_Lichenes_, and _Algæ_ have completely run into one another, and,
when the lowest forms of each are alone considered, even the animal and
vegetable kingdoms cease to have a definite frontier.
If it is permissible to speak of the relations of living forms to one
another metaphorically, the similitude chosen must undoubtedly be that of a
common root, whence two main trunks, one representing the vegetable and one
the animal world, spring; and, each dividing into a few main branches,
these subdivide into multitudes of branchlets and these into smaller groups
of twigs.
As Lamarck has well said--[Footnote: _Philosophie Zoologique_,
première partie, chap. iii.] "Il n'y a que ceux qui se sont longtemps et
fortement occupés de la détermination des espèces, et qui ont consulté de
riches collections, qui peuvent savoir jusqu'à quel point les
_espèces_, parmi les corps vivants se fondent les unes dans les
autres, et qui ont pu se convaincre que, dans les parties où nous voyons
des _espèces_ isolès, cela n'est ainsi que parcequ'il nous en manque
d'autres qui en sont plus voisines et que nous n'avons pas encore
recueillies.
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