Darwiniana : Essays — Volume 02Huxley, Thomas Henry
Science
Darwiniana : Essays — Volume 02
Huxley, Thomas Henry
Darwin, Charles, 1809-1882; Evolution (Biology)
The better its structure is adapted to that of the trap, the more
will it be able to profit by the bait, whether of honey or of pollen, to
the exclusion of its competitors. Thus, by a sort of action and reaction, a
two-fold series of adaptive modifications will be brought about.
In 1865, the important bearing of this subject on his theory led Darwin to
commence a great series of laborious and difficult experiments on the
fertilisation of plants, which occupied him for eleven years, and furnished
him with the unexpectedly strong evidence in favour of the influence of
crossing which he published in 1876, under the title of "The Effects of
Cross and Self Fertilisation in the Vegetable Kingdom." Incidentally, as it
were, to this heavy piece of work, he made the remarkable series of
observations on the different arrangements by which crossing is favoured
and, in many cases, necessitated, which appeared in the work on "The
Different Forms of Flowers in Plants of the same Species" in 1877.
In the course of the twenty years during which Darwin was thus occupied in
opening up new regions of investigation to the botanist and showing the
profound physiological significance of the apparently meaningless
diversities of floral structure, his attention was keenly alive to any
other interesting phenomena of plant life which came in his way. In his
correspondence, he not unfrequently laughs at himself for his ignorance of
systematic botany; and his acquaintance with vegetable anatomy and
physiology was of the slenderest. Nevertheless, if any of the less common
features of plant life came under his notice, that imperious necessity of
seeking for causes which nature had laid upon him, impelled, and indeed
compelled, him to inquire the how and the why of the fact, and its bearing
on his general views. And as, happily, the atavic tendency to frame
hypotheses was accompanied by an equally strong need to test them by
well-devised experiments, and to acquire all possible information before
publishing his results, the effect was that he touched no topic without
elucidating it.
Thus the investigation of the operations of insectivorous plants, embodied
in the work on that topic published in 1875, was started fifteen years
before, by a passing observation made during one of Darwin's rare holidays.
"In the summer of 1860, I was idling and resting near Hartfield, where two
species of Drosera abound; and I noticed that numerous insects had been
entrapped by the leaves. I carried home some plants, and on giving them
some insects saw the movements of the tentacles, and this made me think it
possible that the insects were caught for some special purpose.
Fortunately, a crucial test occurred to me, that of placing a large number
of leaves in various nitrogenous and non-nitrogenous fluids of equal
density; and as soon as I found that the former alone excited energetic
movements, it was obvious that here was a fine new field for
investigation." (I, p. 95.)
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