Darwiniana : Essays — Volume 02Huxley, Thomas Henry
Science
Darwiniana : Essays — Volume 02
Huxley, Thomas Henry
Darwin, Charles, 1809-1882; Evolution (Biology)
These are differences enough in regard to their external appearance; but
these differences are by no means the whole or even the most important of
the differences which obtain between these birds. There is hardly a single
point of their structure which has not become more or less altered; and to
give you an idea of how extensive these alterations are, I have here some
very good skeletons, for which I am indebted to my friend, Mr. Tegetmeier,
a great authority in these matters; by means of which, if you examine them
by and by, you will be able to see the enormous difference in their bony
structures.
I had the privilege, some time ago, of access to some important MSS. of Mr.
Darwin, who, I may tell you, has taken very great pains and spent much
valuable time and attention on the investigation of these variations, and
getting together all the facts that bear upon them. I obtained from these
MSS. the following summary of the differences between the domestic breeds
of pigeons; that is to say, a notification of the various points in which
their organisation differs. In the first place, the back of the skull may
differ a good deal, and the development of the bones of the face may vary a
great deal; the back varies a good deal; the shape of the lower jaw varies;
the tongue varies very greatly, not only in correlation to the length and
size of the beak, but it seems also to have a kind of independent variation
of its own. Then the amount of naked skin round the eyes, and at the base
of the beak, may vary enormously; so may the length of the eyelids, the
shape of the nostrils, and the length of the neck. I have already noticed
the habit of blowing out the gullet, so remarkable in the Pouter, and
comparatively so in the others. There are great differences, too, in the
size of the female and the male, the shape of the body, the number and
width of the processes of the ribs, the development of the ribs, and the
size, shape, and development of the breastbone. We may notice, too--and I
mention the fact because it has been disputed by what is assumed to be high
authority,--the variation in the number of the sacral vertebrae. The number
of these varies from eleven to fourteen, and that without any diminution in
the number of the vertebrae of the back or of the tail. Then the number and
position of the tail-feathers may vary enormously, and so may the number of
the primary and secondary feathers of the wings. Again, the length of the
feet and of the beak,--although they have no relation to each other, yet
appear to go together,--that is, you have a long beak wherever you have
long feet. There are differences also in the periods of the acquirement of
the perfect plumage--the size and shape of the eggs--the nature of flight,
and the powers of flight--so-called _"homing"_ birds having enormous
flying powers; [Footnote: The _"Carrier,"_ I learn from Mr.
Tegetmeier, does not _carry_; a high-bred bird of this breed being but
a poor flier.
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