Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
These useless or non-adaptive characters are, apparently, of the same
nature as the "sports" that arise in our domestic productions, but
which, as Mr. Darwin says, without the aid of selection would soon
disappear; while some of them may be correlations with other characters
which are or have been useful. Some of these correlations are very
curious. Mr. Tegetmeier informed Mr. Darwin that the young of white,
yellow, or dun-coloured pigeons are born almost naked, whereas other
coloured pigeons are born well clothed with down. Now, if this
difference occurred between wild species of different colours, it might
be said that the nakedness of the young could not be of any use. But the
colour with which it is correlated might, as has been shown, be useful
in many ways. The skin and its various appendages, as horns, hoofs,
hair, feathers, and teeth, are homologous parts, and are subject to very
strange correlations of growth. In Paraguay, horses with curled hair
occur, and these always have hoofs exactly like those of a mule, while
the hair of the mane and tail is much shorter than usual. Now, if any
one of these characters were useful, the others correlated with it might
be themselves useless, but would still be tolerably constant because
dependent on a useful organ. So the tusks and the bristles of the boar
are correlated and vary in development together, and the former only may
be useful, or both may be useful in unequal degrees.
The difficulty as to how individual differences or sports can become
fixed and perpetuated, if altogether useless, is evaded by those who
hold that such characters are exceedingly common. Mr. Romanes says that,
upon his theory of physiological selection, "it is quite intelligible
that when a varietal form is differentiated from its parent form by the
bar of sterility, any little meaningless peculiarities of structure or
of instinct _should at first be allowed to arise_, and that they should
then _be allowed to perpetuate themselves_ by heredity," until they are
finally eliminated by disuse. But this is entirely begging the
question. Do meaningless peculiarities, which we admit often arise as
spontaneous variations, ever perpetuate themselves in all the
individuals constituting a variety or race, without selection either
human or natural? Such characters present themselves as unstable
variations, and as such they remain, unless preserved and accumulated by
selection; and they can therefore never become "specific" characters
unless they are strictly correlated with some useful and important
peculiarities.
As bearing upon this question we may refer to what is termed Delboeuf's
law, which has been thus briefly stated by Mr. Murphy in his work on
_Habit and Intelligence_, p. 241.
Public-domain text, read in full here on John Shaqi.
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