Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
During the numerous experiments that have been made on the hybridisation
of plants similar peculiarities have been noticed, some individuals
being capable, others incapable, of being crossed with a distinct
species. The same individual peculiarities are found in varieties,
species, and genera. Kölreuter crossed five varieties of the common
tobacco (Nicotiana tabacum) with a distinct species, Nicotiana
glutinosa, and they all yielded very sterile hybrids; but those raised
from one variety were less sterile, in all the experiments, than the
hybrids from the four other varieties. Again, most of the species of the
genus Nicotiana have been crossed, and freely produce hybrids; but one
species, N. acuminata, not particularly distinct from the others, could
neither fertilise, nor be fertilised by, any of the eight other species
experimented on. Among genera we find some--such as Hippeastrum, Crinum,
Calceolaria, Dianthus--almost all the species of which will fertilise
other species and produce hybrid offspring; while other allied genera,
as Zephyranthes and Silene, notwithstanding the most persevering
efforts, have not produced a single hybrid even between the most closely
allied species.
_Dimorphism and Trimorphism._
Peculiarities in the reproductive system affecting individuals of the
same species reach their maximum in what are called heterostyled, or
dimorphic and trimorphic flowers, the phenomena presented by which form
one of the most remarkable of Mr. Darwin's many discoveries. Our common
cowslip and primrose, as well as many other species of the genus
Primula, have two kinds of flowers in about equal proportions. In one
kind the stamens are short, being situated about the middle of the tube
of the corolla, while the style is long, the globular stigma appearing
just in the centre of the open flower. In the other kind the stamens are
long, appearing in the centre or throat of the flower, while the style
is short, the stigma being situated halfway down the tube at the same
level as the stamens in the other form. These two forms have long been
known to florists as the "pin-eyed" and the "thrum-eyed," but they are
called by Darwin the long-styled and short-styled forms (see woodcut).
[Illustration: FIG. 17.--Primula veris (Cowslip).]
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