Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
But there is another set of plants, the trimorphic, in which the styles
and stamens have each three forms--long, medium, and short, and in these
it is possible to have eighteen different crosses. By an elaborate
series of experiments it was shown that the six legitimate unions--that
is, when a plant was fertilised by pollen from stamens of length
corresponding to that of its style in the two other forms--were all
abundantly fertile; while the twelve illegitimate unions, when a plant
was fertilised by pollen from stamens of a different length from its
own style, in any of the three forms, were either comparatively or
wholly sterile.[52]
We have here a wonderful amount of constitutional difference of the
reproductive organs within a single species, greater than usually occurs
within the numerous distinct species of a genus or group of genera; and
all this diversity appears to have arisen for a purpose which has been
obtained by many other, and apparently simpler, changes of structure or
of function, in other plants. This seems to show us, in the first place,
that variations in the mutual relations of the reproductive organs of
different individuals must be as frequent as structural variations have
been shown to be; and, also, that sterility in itself can be no test of
specific distinctness. But this point will be better considered when we
have further illustrated and discussed the complex phenomena of
hybridity.
_Cases of the Fertility of Hybrids, and of the Infertility of Mongrels._
I now propose to adduce a few cases in which it has been proved, by
experiment, that hybrids between two distinct species are fertile _inter
se_; and then to consider why it is that such cases are so few in
number.
The common domestic goose (Anser ferns) and the Chinese goose (A.
cygnoides) are very distinct species, so distinct that some naturalists
have placed them in different genera; yet they have bred together, and
Mr. Eyton raised from a pair of these hybrids a brood of eight. This
fact was confirmed by Mr. Darwin himself, who raised several fine birds
from a pair of hybrids which were sent him.[53] In India, according to
Mr. Blyth and Captain Hutton, whole flocks of these hybrid geese are
kept in various parts of the country where neither of the pure parent
species exists, and as they are kept for profit they must certainly be
fully fertile.
Another equally striking case is that of the Indian humped and the
common cattle, species which differ osteologically, and also in habits,
form, voice, and constitution, so that they are by no means closely
allied; yet Mr. Darwin assures us that he has received decisive
evidence that the hybrids between these are perfectly fertile _inter
se_.
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