Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
applies to birds, for Darwin was informed by the Rev. W.D. Fox that his
flocks of white and Chinese geese kept distinct.[61]
This constant preference of animals for their like, even in the case of
slightly different varieties of the same species, is evidently a fact
of great importance in considering the origin of species by natural
selection, since it shows us that, so soon as a slight differentiation
of form or colour has been effected, isolation will at once arise by the
selective association of the animals themselves; and thus the great
stumbling-block of "the swamping effects of intercrossing," which has
been so prominently brought forward by many naturalists, will be
completely obviated.
If now we combine with this fact the correlation of colour with
important constitutional peculiarities, and, in some cases, with
infertility; and consider, further, the curious parallelism that has
been shown to exist between the effects of changed conditions and the
intercrossing of varieties in producing either an increase or a decrease
of fertility, we shall have obtained, at all events, a starting-point
for the production of that infertility which is so characteristic a
feature of distinct species when intercrossed. All we need, now, is some
means of increasing or accumulating this initial tendency; and to a
discussion of this problem we will therefore address ourselves.
_The Influence of Natural Selection upon Sterility and Fertility._
It will occur to many persons that, as the infertility or sterility of
incipient species would be useful to them when occupying the same or
adjacent areas, by neutralising the effects of intercrossing, this
infertility might have been increased by the action of natural
selection; and this will be thought the more probable if we admit, as we
have seen reason to do, that variations in fertility occur, perhaps as
frequently as other variations. Mr. Darwin tells us that, at one time,
this appeared to him probable, but he found the problem to be one of
extreme complexity; and he was also influenced against the view by many
considerations which seemed to render such an origin of the sterility or
infertility of species when intercrossed very improbable. The fact that
species which occupy distinct areas, and which nowhere come in contact
with each other, are often sterile when crossed, is one of the
difficulties; but this may perhaps be overcome by the consideration
that, though now isolated, they may, and often must, have been in
contact at their origination. More important is the objection that
natural selection could not possibly have produced the difference that
often occurs between reciprocal crosses, one of these being sometimes
fertile, while the other is sterile. The extremely different amounts of
infertility or sterility between different species of the same genus,
the infertility often bearing no proportion to the difference between
the species crossed, is also an important objection. But none of these
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