Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
Now, let us suppose that a partial sterility of the hybrids between the
two forms arises, in correlation with the different modes of life and
the slight external or internal peculiarities that exist between them,
both of which we have seen to be real causes of infertility. The result
will be that, even if the hybrids between the two forms are still freely
produced, these hybrids will not themselves increase so rapidly as the
two pure forms; and as these latter are, by the terms of the problem,
better suited to their conditions of life than are the hybrids between
them, they will not only increase more rapidly, but will also tend to
supplant the hybrids altogether whenever the struggle for existence
becomes exceptionally severe. Thus, the more complete the sterility of
the hybrids the more rapidly will they die out and leave the two parent
forms pure. Hence it will follow that, if there is greater infertility
between the two forms in one part of the area than the other, these
forms will be kept more pure wherever this greater infertility prevails,
will therefore have an advantage at each recurring period of severe
struggle for existence, and will thus ultimately supplant the less
infertile or completely fertile forms that may exist in other portions
of the area. It thus appears that, in such a case as here supposed,
natural selection would preserve those portions of the two breeds which
were most infertile with each other, or whose hybrid offspring were most
infertile; and would, therefore, if variations in fertility continued to
arise, tend to increase that infertility. It must particularly be noted
that this effect would result, not by the preservation of the infertile
variations on account of their infertility, but by the inferiority of
the hybrid offspring, both as being fewer in numbers, less able to
continue their race, and less adapted to the conditions of existence
than either of the pure forms. It is this inferiority of the hybrid
offspring that is the essential point; and as the number of these
hybrids will be permanently less where the infertility is greatest,
therefore those portions of the two forms in which infertility is
greatest will have the advantage, and will ultimately survive in the
struggle for existence.
Public-domain text, read in full here on John Shaqi.
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