Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
If now in the second and each succeeding year a similar proportion as at
first (10 per cent) of the physiological variety is produced afresh from
the ranks of the normal variety, the same rate of diminution will go on,
and it will be found that, on the most favourable estimate, the
physiological variety can never exceed 12,000 to the 88,000 of the
normal form of the species, as shown by the following table:--
1st Year. 10,000 of physiological variety to 90,000 of normal variety.
2d " 1,220 + 10,000 again produced.
3d " 16 + 1,220 + 10,000 do. = 11,236
4th " O + 16 + 1,220 + 10,000 do. = 11,236
5th " O + 16 + 1,220 + 10,000 = 11,236
and so on for any number of generations.
In the preceding discussion we have given the theory the advantage of
the large proportion of 10 per cent of this very exceptional variety
arising in its midst year by year, and we have seen that, even under
these favourable conditions, it is unable to increase its numbers much
above its starting-point, and that it remains wholly dependent on the
continued renewal of the variety for its existence beyond a few years.
It appears, then, that this form of inter-specific sterility cannot be
increased by natural or any other known form of selection, but that it
contains within itself its own principle of destruction. If it is
proposed to get over the difficulty by postulating a larger percentage
of the variety annually arising within the species, we shall not affect
the law of decrease until we approach equality in the numbers of the two
varieties. But with any such increase of the physiological variety the
species itself would inevitably suffer by the large proportion of
sterile unions in its midst, and would thus be at a great disadvantage
in competition with other species which were fertile throughout. Thus,
natural selection will always tend to weed out any species with too
great a tendency to sterility among its own members, and will therefore
prevent such sterility from becoming the general characteristic of
varying species, which this theory demands should be the case.
On the whole, then, it appears clear that no form of infertility or
sterility between the individuals of a species, can be increased by
natural selection unless correlated with some useful variation, while
all infertility not so correlated has a constant tendency to effect its
own elimination. But the opposite property, fertility, is of vital
importance to every species, and gives the offspring of the individuals
which possess it, in consequence of their superior numbers, a greater
chance of survival in the battle of life. It is, therefore, directly
under the control of natural selection, which acts both by the
self-preservation of fertile and the self-destruction of infertile
stocks--except always where correlated as above, when they become
useful, and therefore subject to be increased by natural selection.
Public-domain text, read in full here on John Shaqi.
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