Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
To many persons it will seem impossible that such beautiful and detailed
resemblances as those now described--and these are only samples of
thousands that occur in all parts of the world--can have been brought
about by the preservation of accidental useful variations. But this will
not seem so surprising if we keep in mind the facts set forth in our
earlier chapters--the rapid multiplication, the severe struggle for
existence, and the constant variability of these and all other
organisms. And, further, we must remember that these delicate
adjustments are the result of a process which has been going on for
millions of years, and that we now see the small percentage of successes
among the myriads of failures. From the very first appearance of insects
and their various kinds of enemies the need of protection arose, and was
usually most easily met by modifications of colour. Hence, we may be
sure that the earliest leaf-eating insects acquired a green colour as
one of the necessities of their existence; and, as the species became
modified and specialised, those feeding on particular species of plants
would rapidly acquire the peculiar tints and markings best adapted to
conceal them upon those plants. Then, every little variation that, once
in a hundred years perhaps, led to the preservation of some larva which
was thereby rather better concealed than its fellows, would form the
starting-point of a further development, leading ultimately to that
perfection of imitation in details which now astonishes us. The
researches of Dr. Weismann illustrate this progressive adaptation. The
very young larvae of several species are green or yellowish without any
markings; they then, in subsequent moults, obtain certain markings, some
of which are often lost again before the larva is fully grown. The early
stages of those species which, like elephant hawk-moths (Chaerocampa),
have the anterior segments elongated and retractile, with large eye-like
spots to imitate the head of a vertebrate, are at first like those of
non-retractile species, the anterior segments being as large as the
rest. After the first moult they become smaller, comparatively; but it
is only after the second moult that the ocelli begin to appear, and
these are not fully defined till after the third moult. This progressive
development of the individual--the ontogeny--gives us a clue to the
ancestral development of the whole race--the phylogeny; and we are
enabled to picture to ourselves the very slow and gradual steps by which
the existing perfect adaptation has been brought about. In many larvae
great variability still exists, and in some there are two or more
distinctly-coloured forms--usually a dark and a light or a brown and a
green form. The larva of the humming-bird hawk-moth (Macroglossa
stellatarum) varies in this manner, and Dr. Weismann raised five
varieties from a batch of eggs from one moth. It feeds on species of
bedstraw (Galium verum and G. mollugo), and as the green forms are less
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