Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
Recognition marks during flight are very important for all birds which
congregate in flocks or which migrate together; and it is essential
that, while being as conspicuous as possible, the marks shall not
interfere with the general protective tints of the species when at rest.
Hence they usually consist of well-contrasted markings on the wings and
tail, which are concealed during repose but become fully visible when
the bird takes flight. Such markings are well seen in our four British
species of shrikes, each having quite different white marks on the
expanded wings and on the tail feathers; and the same is the case with
our three species of Saxicola--the stone-chat, whin-chat, and
wheat-ear--which are thus easily recognisable on the wing, especially
when seen from above, as they would be by stragglers looking out for
their companions. The figures opposite, of the wings of two African
species of stone-curlew which are sometimes found in the same districts,
well illustrates these specific recognition marks. Though not very
greatly different to our eyes, they are no doubt amply so to the sharp
vision of the birds themselves.
Besides the white patches on the primaries here shown, the secondary
feathers are, in some cases, so coloured as to afford very distinctive
markings during flight, as seen in the central secondary quills of two
African coursers (Fig. 21).
[Illustration: FIG. 20.--Oedicnemus vermiculatus (above). Oe.
senegalensis (below).]
Most characteristic of all, however, are the varied markings of the
outer tail-feathers, whose purpose is so well shown by their being
almost always covered during repose by the two middle feathers, which
are themselves quite unmarked and protectively tinted like the rest of
the upper surface of the body. The figures of the expanded tails of two
species of East Asiatic snipe, whose geographical ranges overlap each
other, will serve to illustrate this difference; which is frequently
much greater and modified in an endless variety of ways (Fig. 22).
Numbers of species of pigeons, hawks, finches, warblers, ducks, and
innumerable other birds possess this class of markings; and they
correspond so exactly in general character with those of the mammalia,
already described, that we cannot doubt they serve a similar
purpose.[86]
[Illustration: FIG. 21.--Secondary quills.]
[Illustration: FIG. 22.--Scolopax megala (upper). S. stenura (lower).]
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