When we examine the various modes in which the cross-fertilisation of flowers is brought about, we find that some are comparatively simple in their operation and needful adjustments, others highly complex. The simple methods belong to four principal classes:--(1) By dichogamy--that is, by the anthers and the stigma becoming mature or in a fit state for fertilisation at slightly different times on the same plant. The result of this is that, as plants in different stations, on different soils, or exposed to different aspects flower earlier or later, the mature pollen of one plant can only fertilise some plant exposed to somewhat different conditions or of different constitution, whose stigma will be mature at the same time; and this difference has been shown by Darwin to be that which is adapted to secure the fullest benefit of cross-fertilisation. This occurs in Geranium pratense, Thymus serpyllum, Arum maculatum, and many others. (2) By the flower being self-sterile with its own pollen, as in the crimson flax. This absolutely prevents self-fertilisation. (3) By the stamens and anthers being so placed that the pollen cannot fall upon the stigma, while it does fall upon a visiting insect which carries it to the stigma of another flower. This effect is produced in a variety of very simple ways, and is often aided by the motion of the stamens which bend down out of the way of the stigmas before the pollen is ripe, as in Malva sylvestris (see Fig. 28). (4) By the male and female flowers being on different plants, forming the class Dioecia of Linnaeus. In these cases the pollen may be carried to the stigmas either by the wind or by the agency of insects. [Illustration: FIG. 28. Malva sylvestris, adapted for insect-fertilisation. Malva rotundifolia, adapted for self-fertilisation.] Now these four methods are all apparently very simple, and easily produced by variation and selection. They are applicable to flowers of any shape, requiring only such size and colour as to attract insects, and some secretion of nectar to ensure their repeated visits, characters common to the great majority of flowers. All these methods are common, except perhaps the second; but there are many flowers in which the pollen from another plant is prepotent over the pollen from fertilisation, the same flower, and this has nearly the same effect as self-sterility if the flowers are frequently crossed by insects. We cannot help asking, therefore, why have other and much more elaborate methods been needed? And how have the more complex arrangements of so many flowers been brought about? Before attempting to answer these questions, and in order that the reader may appreciate the difficulty of the problem and the nature of the facts to be explained, it will be necessary to give a summary of the more elaborate modes of securing cross-fertilisation. (1) We first have dimorphism and heteromorphism, the phenomena of which have been already sketched in our seventh chapter.