Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
A French botanist, Mons. A. Jordan, has collected numerous forms of a
common little plant, the spring whitlow-grass (Draba verna); he has
cultivated these for several successive years, and declares that they
preserve their peculiarities unchanged; he also says that they each come
true from seed, and thus possess all the characteristics of true
species. He has described no less than fifty-two such species or
permanent varieties, all found in the south of France; and he urges
botanists to follow his example in collecting, describing, and
cultivating all such varieties as may occur in their respective
districts. Now, as the plant is very common almost all over Europe and
ranges from North America to the Himalayas, the number of similar forms
over this wide area would probably have to be reckoned by hundreds if
not by thousands.
The class of facts now adduced must certainly be held to prove that in
many large genera and in some single species there is a very large
amount of variation, which renders it quite impossible for experts to
agree upon the limits of species. We will now adduce a few striking
cases of individual variation.
The distinguished botanist, Alp. de Candolle, made a special study of
the oaks of the whole world, and has stated some remarkable facts as to
their variability. He declares that on the same branch of oak he has
noted the following variations: (1) In the length of the petiole, as one
to three; (2) in the form of the leaf, being either elliptical or
obovoid; (3) in the margin being entire, or notched, or even pinnatifid;
(4) in the extremity being acute or blunt; (5) in the base being sharp,
blunt, or cordate; (6) in the surface being pubescent or smooth; (7) the
perianth varies in depth and lobing; (8) the stamens vary in number,
independently; (9) the anthers are mucronate or blunt; (10) the fruit
stalks vary greatly in length, often as one to three; (11) the number of
fruits varies; (12) the form of the base of the cup varies; (13) the
scales of the cup vary in form; (14) the proportions of the acorns vary;
(15) the times of the acorns ripening and falling vary.
Besides this, many species exhibit well-marked varieties which have been
described and named, and these are most numerous in the best-known
species. Our British oak (Quercus robur) has twenty-eight varieties;
Quercus Lusitanica has eleven; Quercus calliprinos has ten; and Quercus
coccifera eight.
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