Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
But the most curious and striking confirmation of this portion of Mr.
Darwin's theory is afforded by the case of Kerguelen Island. This island
was visited by the _Transit of Venus_ expedition. It is one of the
stormiest places on the globe, being subject to almost perpetual gales,
while, there being no wood, it is almost entirely without shelter. The
Rev. A.E. Eaton, an experienced entomologist, was naturalist to the
expedition, and he assiduously collected the few insects that were to be
found. All were incapable of flight, and most of them entirely without
wings. They included a moth, several flies, and numerous beetles. As
these insects could hardly have reached the islands in a wingless state,
even if there were any other known land inhabited by them--which there
is not--we must assume that, like the Madeiran insects, they were
originally winged, and lost their power of flight because its possession
was injurious to them.
It is no doubt due to the same cause that some butterflies on small and
exposed islands have their wings reduced in size, as is strikingly the
case with the small tortoise-shell butterfly (Vanessa urticae)
inhabiting the Isle of Man, which is only about half the size of the
same species in England or Ireland; and Mr. Wollaston notes that Vanessa
callirhoe--a closely allied South European form of our red-admiral
butterfly--is permanently smaller in the small and bare island of Porto
Santo than in the larger and more wooded adjacent island of Madeira.
A very good example of comparatively recent divergence of character, in
accordance with new conditions of life, is afforded by our red grouse.
This bird, the Lagopus scoticus of naturalists, is entirely confined to
the British Isles. It is, however, very closely allied to the willow
grouse (Lagopus albus), a bird which ranges all over Europe, Northern
Asia, and North America, but which, unlike our species, changes to white
in winter. No difference in form or structure can be detected between
the two birds, but as they differ so decidedly in colour--our species
being usually rather darker in winter than in summer, while there are
also slight differences in the call-note and in habits,--the two species
are generally considered to be distinct. The differences, however, are
so clearly adaptations to changed conditions that we can hardly doubt
that, during the early part of the glacial period, when our islands were
united to the continent, our grouse was identical with that of the rest
of Europe. But when the cold passed away and our islands became
permanently separated from the mainland, with a mild and equable climate
and very little snow in winter, the change to white at that season
became hurtful, rendering the birds more conspicuous instead of serving
as a means of concealment. The colour was, therefore, gradually changed
by the process of variation and natural selection; and as the birds
obtained ample shelter among the heather which clothes so many of our
Public-domain text, read in full here on John Shaqi.
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