Discourses: Biological & Geological: EssaysHuxley, Thomas Henry
Science
Discourses: Biological & Geological: Essays
Huxley, Thomas Henry
Biology; Geology
But when we turn to the higher _Vertebrata_, the results of recent
investigations, however we may sift and criticise them, seem to me to
leave a clear balance in favour of the doctrine of the evolution of
living forms one from another. Nevertheless, in discussing this question,
it is very necessary to discriminate carefully between the different
kinds of evidence from fossil remains which are brought forward in favour
of evolution.
Every fossil which takes an intermediate place between forms of life
already known, may be said, so far as it is intermediate, to be evidence
in favour of evolution, inasmuch as it shows a possible road by which
evolution may have taken place. But the mere discovery of such a form
does not, in itself, prove that evolution took place by and through it,
nor does it constitute more than presumptive evidence in favour of
evolution in general. Suppose A, B, C to be three forms, while B is
intermediate in structure between A and C. Then the doctrine of evolution
offers four possible alternatives. A may have become C by way of B; or C
may have become A by way of B; or A and C may be independent
modifications of B; or A, B, and C may be independent modifications of
some unknown D. Take the case of the Pigs, the _Anoplothcridoe_, and the
Ruminants. The _Anoplothcridoe_ are intermediate between the first and
the last; but this does not tell us whether the Ruminants have come from
the Pigs, or the Pigs from Ruminants, or both from _Anoplothcridoe_, or
whether Pigs, Ruminants, and _Anoplotlicridoe_ alike may not have
diverged from some common stock.
But if it can be shown that A, B, and C exhibit successive stages in the
degree of modification, or specialisation, of the same type; and if,
further, it can be proved that they occur in successively newer deposits,
A being in the oldest and C in the newest, then the intermediate
character of B has quite another importance, and I should accept it,
without hesitation, as a link in the genealogy of C. I should consider
the burden of proof to be thrown upon any one who denied C to have been
derived from A by way of B, or in some closely analogous fashion; for it
is always probable that one may not hit upon the exact line of filiation,
and, in dealing with fossils, may mistake uncles and nephews for fathers
and sons.
I think it necessary to distinguish between the former and the latter
classes of intermediate forms, as _intercalary types_ and _linear types_.
When I apply the former term, I merely mean to say that as a matter of
fact, the form B, so named, is intermediate between the others, in the
sense in which the _Anoplotherium_ is intermediate between the Pigs and
the Ruminants--without either affirming, or denying, any direct genetic
relation between the three forms involved. When I apply the latter term,
on the other hand, I mean to express the opinion that the forms A, B, and
C constitute a line of descent, and that B is thus part of the lineage of
C.
Public-domain text, read in full here on John Shaqi.
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